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Evolution PCB4674 Midterm exam2 Name:__________________________________ Mar 22 2005 ID:________________ For each multiple choice question select the single best answer. Answer questions 1 to 20 on your scantron sheet. Answer the remaining questions in the space provided or on the figures. Brevity is rewarded. Please print clearly. Each multiple choice question is worth 1 point. For short answer questions, the number of points is indicated in parentheses at the end of each question. 1 a) b) c) d) e) A point mutation is: a single allele substitution in DNA a single base substitution in DNA a single chromosome substitution in DNA a single gene substitution in DNA none of the above b 2 What is the Red Queen hypothesis? a) sexual selection is driven by female choice b) organisms have to evolve as fast as they can just to keep pace with evolution of other organisms around them c) genetic diversity is maintained by natural selection d) allele frequencies will not change generation after generation e) none of the above b 3 a) b) c) d) e) An insertion or deletion of a base changes the codon reading sequence. This is knows as: frameshift mutation point mutation chromosome inversion transversion none of the above a 4 Given a low degree of heritability for a trait and a low selection differential, what would be the predicted evolutionary response to selection? R=h2S a) high b) low c) intermediate d) any of the above e) insufficient information to answer that question b 5 The vast majority of mutations are: a) strongly beneficial b) c) d) e) neutral to slightly beneficial neutral to slightly deleterious strongly deleterious lethal a) b) c) d) e) Genetic drift is most important in: large populations small populations asexual populations sexually reproducing populations all of the above a) b) c) d) e) Random genetic drift will cause the average heterozygosity in a population to: increase decrease remain stable all of the above are possible none of the above c 6 b 7 b 8 If 2 groups compete for the same ecological niche, one will drive the other to extinction. This is known as: a) character displacement b) community interaction c) competitive exclusion d) resource partitioning e) none of the above c 9 a) b) c) d) e) Evolution above the species level is known as: microevolution gradualism population genetics saltation none of the above e 10 The rapid diversification of a small group of ancestral species into a large number of descendant species that occupy a wide range of ecological niches is known as: a) adaptive radiation b) gradualism c) migration d) drift e) sexual selection a 11 Evolutionary changes that fluctuate around a mean value and give the overall appearance of stasis are known as: a) punctuated equilibrium b) gradualism c) selection d) mutation e) zigzag evolution e 12 a) b) c) d) e) What is a polymorphism? a difference in the rate of evolutionary change expression of different protein products from the same allele multiple mutations in a single individual more than 1 possible allele at a particular locus none of the above a) b) c) d) e) What effect does inbreeding have on the frequency of homozygotes in a population? increases homozygotes decreases homozygotes stabilizes frequency of homozygotes all of the above are possible none of the above a) b) c) d) e) In the realm of population genetics, evolution is defined as: any violation of Hardy-Weinberg Equilibrium assumptions the appearance of a new species a change in allele frequencies across generations all of the above population genetics has nothing to do with evolution a) b) c) d) e) Which of the following are mechanisms of divergence that can lead to speciation? genetic drift sexual selection natural selection all of the above none of the above d 13 a 14 c 15 d 16 Define resource partitioning. (1) Species minimize harmful effects of direct competition by using different aspects of their common environment 17 List 2 assumptions required for Hardy-Weinberg Equilibrium. The requisite assumptions for HWE are unrealistic and unlikely to be found in any population so what is the value of this model? (3) No selection, no mutation, no migration , no drift, no assortative mating Allows us to quantify what will happen when each assumption is violated 18 Briefly describe Eldredge and Gould’s concept of punctuated equilibrium. What derogatory term was applied to their concept? (3) Evolution tends to be characterized by long periods of stasis punctuated by episodes of very fast development of new forms Evolution by jerks 19 Hardy-Weinberg equilibrium states that regardless of allele frequencies, genotype frequencies will go to and remain at: (1) p2 + 2pq + q2 = 1 20 Distinguish between allopatric and sympatric speciation. Give an example (real or hypothetical) of allopatric speciation. (3) allopatric speciation – populations evolve in geographic isolation sympatric speciation – populations evolve in geographic proximity 21 Distinguish between Batesian and Muellerian mimicry. (3) Batesian – 1 species mimics another Muellerian – 2 species share similar aposematic coloration 22 Distinguish between coevolution and cospeciation. (3) coevolution – evolutionary changes in 1+ species in response to changes in other species in the same community cospeciation – speciation process that occurs in 2 interacting species simultaneously 23 Label these selection modes. (3) directional 24 disruptive stabilizing Label these two hypotheses for the evolution of different morphologies over time. (2) saltation gradualism 25 For a population of mice, the mean midparent value for tail length for the entire population =18.3cm and mean midparent value for the one third of the population with the longest tails =19.9cm. If the mean midoffspring value for the entire population =18.7 and the mean midoffspring value for the offspring of the parents with the longest tails =20.1, what is the heritability of the selected trait? Given a selection differential of 0.8, what is the predicted evolutionary response (R)? (3) O * -O R = P * -P S heritability (h2) = 0.875 selection differential (R) = 0.7 26 You observe a field of wildflowers with the following genotype frequencies: AA: 144, Aa: 100, aa: 81. i) What are the allele frequencies? ii) What are the expected genotype frequencies under Hardy-Weinberg equilibrium assumptions? iii) Is this population in HardyWeinberg equilibrium (ie. is the observed genotype frequency close to the expected frequency)? (6) i) A: 0.5969; a: 0.4031 ii) AA: 0.3563; Aa: 0.4812; aa: 0.1625 iii) no, observed numbers are very different from expected 27 Given initial allele frequencies of A: 0.9 and a: 0.1, calculate the i) genotype frequencies for a population of 1000 individuals ii) genotype frequencies after selection eliminates 40% of all homozygotes and ii) final allele frequencies. Which conclusion(s) is violated. (6) i) AA: 810; Aa: 180: aa: 10 ii) AA: 486; Aa: 180; aa: 6 iii) A: 0.8571; a: 0.1429 selection violates conclusion 1: allele frequencies in a population will not change, generation after generation 28 Choose any one of the required chapters from The Origin of Species (Difficulties on theory, Instinct, Hybridism, Geological record, Geological succession) and provide a brief summary of the chapter including at least 2 examples mentioned by Darwin. (8)