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Nervous System THE BRAIN CONTINUED Review White versus grey matter Ventricles 4 brain regions 4 lobes of cerebral hemispheres 3 layers of cerebrum Cortex Motor Sensory Association White matter tracts Gray matter Brain Regions 4 Adult brain regions 1. Cerebral hemispheres (cerebrum) 2. Diencephalon 3. 4. Cerebellum Brain stem (midbrain, pons, and medulla) Diencephalon Three paired structures Thalamus Hypothalamus Epithalamus Encloses the third ventricle Cerebral hemisphere Septum pellucidum Interthalamic adhesion (intermediate mass of thalamus) Interventricular foramen Anterior commissure Hypothalamus Optic chiasma Pituitary gland Mammillary body Pons Medulla oblongata Corpus callosum Fornix Choroid plexus Thalamus (encloses third ventricle) Posterior commissure Pineal gland (part of epithalamus) Corpora quadrigemina MidCerebral brain aqueduct Arbor vitae (of cerebellum) Fourth ventricle Choroid plexus Cerebellum Spinal cord Figure 12.12 Diencephalon Thalamus Several nuclei Gateway of the cerebral cortex Major relay station for most sensory impulses Diencephalon Thalamus Relay center for cerebral activation Associated with reticular formation Relay center for somatosensory information Except olfaction Coma is associated with thalamic injury Vegetative state = damage to cortical pathways Refer to diagram on CNS 8 Diencephalon Hypothalamus Inferior to the thalamus Forms portions of walls of the third ventricle Consists of a number of nuclei Diencephalon Hypothalamus Infundibulum Mammillary bodies Responsible for most neurogenic homeostasis of the body Diencephalon Hypothalamic function Autonomic control center for many visceral functions Examples Blood pressure, rate and force of heartbeat Regulates body temperature Hunger and G.I tract regulation Center for emotional response Example Tactile sexual response Diencephalon Hypothalamic Function Water balance and thirst Controls release of hormones by the anterior pituitary and produces posterior pituitary hormones Regulation of sleep-wake cycles Diencephalon Epithalamus Forms roof of third ventricle Pineal gland, choroid plexus Melatonin We’ll discuss it’s endocrine function later…. Brain Regions 4 Adult brain regions 2. Cerebral hemispheres (cerebrum) Diencephalon 3. Brain stem (midbrain, pons and medulla) 4. Cerebellum 1. The Brain Stem Functions Supports most of basic life functions Pathway for fiber tracts Origin for most cranial nerves The Brain Stem Midbrain Associated with visual and auditory reflexes Pupillary reflex Cranial nerves III and IV Red nucleus Descending motor pathways involved in voluntary movement Frontal lobe Olfactory bulb (synapse point of cranial nerve I) Optic chiasma Optic nerve (II) Optic tract Mammillary body Midbrain Pons Temporal lobe Medulla oblongata Cerebellum Spinal cord Figure 12.14 View (a) Optic chiasma Optic nerve (II) Crus cerebri of cerebral peduncles (midbrain) Diencephalon • Thalamus • Hypothalamus Mammillary body Thalamus Hypothalamus Diencephalon Midbrain Oculomotor nerve (III) Trochlear nerve (IV) Pons Brainstem Medulla oblongata Trigeminal nerve (V) Pons Facial nerve (VII) Middle cerebellar peduncle Abducens nerve (VI) Vestibulocochlear nerve (VIII) Pyramid Glossopharyngeal nerve (IX) Hypoglossal nerve (XII) Vagus nerve (X) Ventral root of first cervical nerve Decussation of pyramids Accessory nerve (XI) Spinal cord (a) Ventral view Figure 12.15a Crus cerebri of cerebral peduncles (midbrain) Thalamus View (b) Infundibulum Pituitary gland Superior colliculus Inferior colliculus Trochlear nerve (IV) Trigeminal nerve (V) Pons Superior cerebellar peduncle Middle cerebellar peduncle Facial nerve (VII) Abducens nerve (VI) Glossopharyngeal nerve (IX) Hypoglossal nerve (XII) Inferior cerebellar peduncle Vestibulocochlear nerve (VIII) Olive Thalamus Vagus nerve (X) Hypothalamus Diencephalon Midbrain Accessory nerve (XI) Pons Brainstem Medulla oblongata (b) Left lateral view Figure 12.15b The Brain Stem Pons Bridge between midbrain and medulla oblongata Connects cerebellum to cerebrum and spinal cord Cranial nerves V- VIII (vestibular branch) Crus cerebri of cerebral peduncles (midbrain) Thalamus View (b) Infundibulum Pituitary gland Superior colliculus Inferior colliculus Trochlear nerve (IV) Trigeminal nerve (V) Pons Superior cerebellar peduncle Middle cerebellar peduncle Facial nerve (VII) Abducens nerve (VI) Glossopharyngeal nerve (IX) Hypoglossal nerve (XII) Inferior cerebellar peduncle Vestibulocochlear nerve (VIII) Olive Thalamus Vagus nerve (X) Hypothalamus Diencephalon Midbrain Accessory nerve (XI) Pons Brainstem Medulla oblongata (b) Left lateral view Figure 12.15b The Brain Stem Medulla Continuous with spinal cord Passage of motor & sensory impulses between brain & spinal cord Decussation of tracts in pyramids Longitudinal fissure Lateral ventricle Superior Commissural fibers (corpus callosum) Association fibers Basal nuclei • Caudate • Putamen • Globus pallidus Corona radiata Thalamus Internal capsule Fornix Gray matter Third ventricle White matter Pons Projection fibers Medulla oblongata (a) Decussation of pyramids Figure 12.10a The Brain Center Medulla Vital centers Cardiac Vasomotor Respiratory Swallowing Vomiting Cranial nerves VIII (cochlear branch) -XII Brain Regions 4 Adult brain regions 3. Cerebral hemispheres (cerebrum) Diencephalon Brain stem (midbrain, pons, and medulla) 4. Cerebellum 1. 2. The Cerebellum Dorsal to the pons & medulla Subconsciously provides precise timing & appropriate patterns of skeletal muscle contraction Contains both white & gray matter Anterior lobe Cerebellar cortex Arbor vitae Cerebellar peduncles • Superior • Middle • Inferior Medulla oblongata (b) Flocculonodular lobe Posterior lobe Choroid plexus of fourth ventricle Figure 12.17b The Cerebellum Functions Proprioception Prime mover inhibition and antagonist activation Dysmetria Progression Dysarthria Functional Brain Systems Networks of neurons that work together & span wide areas of the brain Limbic system Reticular formation Functional Brain Systems Limbic system Structures on the medial aspects of cerebral hemispheres and diencephalon Includes parts of the diencephalon and some cerebral structures that encircle the brain stem Septum pellucidum Diencephalic structures of the limbic system •Anterior thalamic nuclei (flanking 3rd ventricle) •Hypothalamus •Mammillary body Olfactory bulb Corpus callosum Fiber tracts connecting limbic system structures •Fornix •Anterior commissure Cerebral structures of the limbic system •Cingulate gyrus •Septal nuclei •Amygdala •Hippocampus •Dentate gyrus •Parahippocampal gyrus Figure 12.18 Functional Brain Systems Limbic system Emotional brain Recognizes angry or fearful facial expressions Assesses danger & elicits the fear response Plays a role in expressing emotions via gestures and resolves mental conflict Connection to pre-frontal cortex allows us to “count to ten” Puts emotional responses to odors Example: skunks = smell bad Functional Brain Systems Reticular formation Broad columns of nuclei along the length of the brain stem Far-flung axonal connections with hypothalamus, thalamus, cerebral cortex, cerebellum & spinal cord Radiations to cerebral cortex Visual impulses Auditory impulses Reticular formation Ascending general sensory tracts (touch, pain, temperature) Descending motor projections to spinal cord Figure 12.19 Functional Brain Systems Functions of the reticular formation 1. Somatic motor control Reticulospinal tract = improves smoothness of movement Autonomic control 2. Respiratory and cardiovascular centers Arousal 3. RAS = filter Pain modulation 4. Can block pain transmission Questions? Homework due in Lab PreLab #2 HW #2 – Synapses (page 1 and 2 in HW section)