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Pitt-London Workshop: Abstracts in alphabetical order Karen Arnold, Putting Anthropomorphism in Context At the end of his 1993 book, The Disorder of Things, John Dupré suggests that science, as a human product, should be evaluated on the basis of its contributions “to the thriving of the sentient beings in this universe” (Dupré 1993, p.264). This paper presents a case study of this type of evaluation by analyzing critiques of evolutionary explanations of rape. Since the late 70’s several sociobiologists and evolutionary psychologists have attempted to provide evolutionary accounts of rape. These accounts have come under attack for their use of the term ‘rape’ to describe both human and animal behavior. Critics have objected to this descriptive vocabulary as illegitimately anthropomorphic. This paper provides a taxonomy of these attacks. With this taxonomy in place, the focus will narrow to an examination of those criticisms that rely on contextual values—those values that “belong to the social and cultural context in which science is done” (Longino 1998, 245). I argue that this anthropomorphic use of language can be legitimately attacked on the grounds that it has the potential for a significantly detrimental impact on society. James Bogen, What we Talk About When we Talk about Causality: Mechanistic explanations depend on descriptions of activities, rather than counterfactual regularities. Systematic Dependence (SD) and Mechanistic accounts of the causality are the most promising alternatives to covering law accounts of causality. Both maintain that to fully understand how an effect was caused would be to understand how it results from the workings of a system of causally relevant items. Both maintain that to understand the influence of a particular part of the system would be to understand its contribution to the system's production of the effect. For Mechanism this is a matter of understanding the activities the item of interest engages in, and what results from them. For SD it is a matter of discerning relations of systematic dependency between the item of interest and the effect to be explained. Systematic dependence, in turn, is to be understood in terms of counterfactual regularities of a special kind between manipulations (also of a special kind) of the "values" (i.e., features, quantitative or qualitative states, etc.) of the item of interest, and the results they produce. According to SD, it is these counterfactual regularities (as opposed to activities) which distinguish non-causal from causally productive relations among things. According to Mechanism, any regularities involved in the operation of the system that produces the effect of interest are to be explained by appeal to the activities. This paper argues that SD cannot show us how to understand causality without activities---partly because the counterfactuals it's account of systematic depends upon are highly problematic. Ingo Brigandt, The "role" a concept plays in science - The case of homology The paper tries to shed some light on the idea that a concept plays a certain role for a scientific field or a research program by using the homology concept as a case study. It is in particular discussed in what sense homology plays a different role for systematics, developmental biology, and molecular biology. Joe Cain, Extinction concepts put to work in the philosophy of biology In the analysis of biological processes, extinction disappears somewhere in the miscellany between the purpose of zoos and teleological progress. That should change. In this paper I undertake a conceptual analysis of extinction and indicate some ways this subject can speak to general themes in the philosophy of science. Conceptual analysis focuses on clarification of extinction definitions, the distinction between normal and mass extinction, and conflicting demands in notions of mass extinction. Relevance to general themes is examined in a small study of causal language used in extinction descriptions. A proposed distinction between natural and artificial extinction is used to consider the conjunction of extinction and the ethics of human impact. Paul Griffiths, Evo-Devo meets the Mind: Towards a Developmental Evolutionary Psychology The view of the evolutionary process presumed in contemporary Evolutionary Psychology is in stark contrast to the views being explored by evolutionary developmental biologists. The fertilised egg is held to contain a program for the adult phenotype, the content of which can be inferred by optimality modeling and knowledge of the ancestral environment. The psychological phenotype is thought to be modular, but this modular structure reflects the fact that the environment contains separate adaptive problems and it too can be discovered by Œadaptive thinking‚. Psychological traits are defined by their adaptive function and the concept of homology is rarely utilized. There is, however, an alternative tradition of research in animal behavior, represented by the work of, amongst others, Patrick Bateson, Peter Klopfer and Gilbert Gottlieb, which focuses evolutionary thinking on the developmental structure of the organism. Evolution is seen as creating systems with a range of developmental potentials, rather than programs for phenotypes. The developmental structure of the organism is accepted as an important determinant of evolution and answers to developmental and adaptive questions are thought to be mutually constraining. In this paper I compare this developmentalist tradition in the study of behavior to evolutionary developmental biology, with the aim of enriching the former with some of the new insights that have emerged in recent years from the study of morphological evolution. I offer some tentative suggestions about how an evolutionary psychology might develop along the lines suggested by this synthesis. Catherine Kendig, Reconstructing the Concept of Homology for Genomics Homology has been one of, if not the most, fecund concepts which has been used towards the understanding of the genomes of the model organisms. The evidence for this claim can be supported best with an examination of current research in comparative genomics. In comparative genomics, the information of genes or segments of the genome, and their location and sequence, are used to search for genes similar to them, known as ‘homologues’. Homologues can be either within that same organism (paralogues), or among different species (orthologues). The importance in finding homologous genes within organisms or across species is that these similarities indicate the possibility of ascribing functions, mechanisms or structures which are required by a variety of species which present the same homology. The interest in structures and functions of genes and proteins common to multiple species is one of the main foci of comparative genomics. Because of this, research into the conservation of genes has been the basis of comparison with regards to homologous genes among diverse organisms. Different causal processes are involved in genetic pathways and mechanisms. Explanations of these depend upon which pathway, structure or mechanism is picked out. Each process has a different causal network to which different explanations refer. What comparative genomics explains are the different causal mechanisms which occur in processes such as differentiation, protein synthesis, and gene regulation. How these processes interact within the organism can only be understood when compared with organisms which possess homologous genes, gene sequences, similar developmental mechanisms, or those whose mechanisms for gene regulation are similar. Explanations which result from comparative genomics contribute to a more comprehensive understanding of both the complex structures and the diverse functions within the genomes of different organisms. There are two related problems which have plagued attempts to define the concept of homology. The first problem arises in clarifying what kind of similarity is involved in a homological comparison. A second problem occurs if more than one concept of homology is needed to pick out the kinds of similarity in different contexts of homological comparison. Homology is usually understood as picking out what counts as ‘the same’ between two or more organisms. Many of the attempts which have been made to define the concept of homology focus on which criteria are used to restrict the kinds of similarity which exist between two or more organisms or parts being compared. These are criteria which can be used reliably to infer shared ancestry. However, there have been many different attempts to define similarity which have produced a profusion of homology concepts. This profusion has led both to the conflation of what counts as ‘the same’ in different contexts and has also muddled the relations of comparison which various concepts use to identify homologues. James Lennox, History and Philosophy of Science: A Phylogenetic Approach In the aftermath of Thomas Kuhn’s The Structure of Scientific Revolutions, there was much discussion about the relationship between the history of science and the philosophy of science. A wider issue was at stake in these discussions: ‘normativism versus ‘naturalism’ in epistemology. If the history of science, at best, gives us reliable information about what actually occurred historically, how can it inform debates about such things as confirmation or explanation in philosophy of science? This essay makes a case for the centrality of historical investigation in the philosophy of science. I will defend what I term the ‘phylogenetic’ approach to the philosophy of science. I will argue that since the foundations and dominant methods of a particular scientific field are shaped by its history, studying that history can give us considerable insight into conceptual and methodological problems in a particular science. The case will be made both on general, philosophical grounds, and by instantiation. I will outline the ways in which looking at the historical origins of the mis-named ‘tautology’ objection to evolutionary theory can be of value in understanding, and dealing with, the problem. Alan C. Love, Evolutionary Morphology and the Integration of Evolution and Development One foundational question in contemporary biology is how to integrate evolution and development. The emerging synthesis (evolutionary developmental biology or “evo-devo”) requires a meshing of disciplines, concepts, and explanatory assumptions (among other things) that have been developed largely in independence over the past century. The nature of the hoped for synthesis is not wholly agreed upon due to divergent viewpoints resulting from this disciplinary independence and, consequently, the mechanics for accomplishing the task are not clearly specified. This paper utilizes historical investigation for philosophical purposes in the way articulated by Jim Lennox in his presentation on the phylogenetic approach to history and philosophy of science in order to explore the question of integrating evolutionary and developmental biology. In the attempt to comprehend the present separation between evolution and development much attention has been paid to the split between genetics and embryology in the early part of the century with its codification in the exclusion of embryology from the Modern Synthesis. This encourages a characterization of “evo-devo” as the integration of developmental genetics with Neo-Darwinism. But there is a largely untold story about the significance of morphology and comparative anatomy (also minimized in the Modern Synthesis) in contemporary discussions. I will attempt to reconstruct part of this story, focusing on the rebirth of functional (and evolutionary) morphology after the 1950's. Functional morphology is critical for understanding the development of a concept central to “evodevo”, evolutionary innovation. Understanding the story about morphology and innovation reveals a different conception of the foundational problem, providing an alternative way of conceptualizing the “evo” and the “devo” to be integrated. (This paper is the beginning of a larger dissertation project on explanations of the origin of evolutionary novelty since the Modern Synthesis with special reference to evolutionary developmental biology.) Peter Machamer and Jacqueline Sullivan, Leveling Reduction Philosophical models of reduction presuppose that there are distinct levels entities (epistemic or ontic) in the world, and that somehow it is desirable to have explanations that are uni-level descriptions. In this paper we argue that explanations, typically, if not always, are multileveled descriptions. One reason for this is that activities are crucial to explanations, and activities typically, are involved with entities at more than one level. We show this by a general argument and in detail by looking at the case of LTP and memory in neuroscience. Two interesting conclusions follow. First, if "level" is used in a the way scientists use it, it is a harmless, adequately clear notion relativized to a research project; neither of these is true when philosophers use "level" in an ontic or episetemic sense. Second, what scientists seem to mean when they talk about "reduction" is finding the mechanisms by which things work, and this is what philosophers ought to mean too. In which case "reduction" becomes a harmless, mostly clear concept that means filling in mechanism schemata. Philosophical talk about reduction and levels obscures the way in which explanations (mechanisms) are sought and given. Gianmatteo Mameli, On The Gene-Centric View of Natural Selection The gene-centric view of natural selection is the view that when natural selection acts upon a population of organisms it always produces changes in the gene pool of that population. There are three distinct arguments that can be or have been proposed in support of such a view: "the argument from development", "the argument from transgenerational stability of form", and "the argument from transgenerational stability of variation". Both the supporters and the critics of the gene-centric view show little or no awareness of the differences between the three arguments. I spell out the three arguments, show how they differ, and show what is wrong with each of them. Sandra Mitchell, Anthropomorphism: Cross-Species Modeling There has been a recent resurgence of interest in anthropomorphism, attributable to both the rise of cognitive ethology and the requirements of various forms of expanded, environmental ethics. The manner and degree to which non-human animals are similar to human beings has thus become a focus of scientific research and a necessary component to our decisions to act morally. At its basis, anthropomorphism involves claims about the similarity of non-human objects or beings to humans. Critics of anthropomorphism often attack the presumptive character of such claims. In this paper I consider a range of stances toward anthropomorphism from global rejections to specific models. The bumper sticker version of this talk could be: science made too easy is bound to be wrong. In the end I will argue that specific anthropomorphic theses are supported or not supported by the same rigorous experimental and logical reasoning as any other scientific model. David Papineau, The Evolution of Means-End Cognition: Why Animals Ain't Smart Why is there a cognitive gulf between other animals and humans? Current fashion favours our greater understanding of Theory of Mind as an answer, and Language is another obvious candidate. But I think that analysis of the evolution of means-end cognitive mechanisms suggests that there may be a further significant difference: where animals will only perform those means which they (or their ancestors) have previously used as a route to some end, humans can employ observation to learn that some novel means is a route to a desired end. In short, human can learn from observation, where animals can learn only from first-hand experience. Karola Stotz, Bringing Life back to Mind: Bodily experience, situated activity, and environmental embeddedness of cognition The center of qualitative experience as cognition, feeling, and awareness, is not the mind alone but the whole organism including beside the mind its body and its environment. And it is the whole organism that, as the center of agency, is, in Kauffman’s words, acting on its own behalf. Living a life instead of computing it allows the organism to act spontaneously and appropriately, an end less achieved by means of rich but passive inner models of the world than by the ‘cheap and efficient production of real-world action in a real-world context’ (Clark). In this paper I shall discuss some of the artificial mysteries of the mind created by mainstream representationalist approaches of cognition, and I will explore some of the implications of reaching beyond the information processing and computation metaphor in cognitive science. D.M.Walsh, The Causes of Adaptation and the Unity of Biology (draft of paper available) Evolutionary Biology has two principal explananda, fit and diversity (Lewontin 1978). Natural selection theory stakes its claim to being the central unifying concept in biology on the grounds that it demonstrates both phenomena to be the consequence of a single process. By now the standard story hardly needs reiterating: Natural selection is a force that operates over a population, preserving the better fit, culling the less fit, and along the way promoting novel solutions to adaptive problems. Amundson’s historical survey of the concept of adaptation captures the idea succinctly: The phenomenon of adaptation is at the core of modern evolutionary biology. Natural selection, the mechanism universally regarded as the primary causal influence on phenotypic evolutionary change, is first and foremost an explanation of adaptation. (1996: 11) At the same time, it appears that the capacity of natural selection to cause adaptations cannot account for every feature of the nature and distribution of biological form. Reason to believe this devolves from the simple fact that the bearers of biological form are organisms and a salient fact about organisms is that each faces the tribunal of the environment as a corporate entity, not as a loose aggregate of independent traits. One consequence of this fact is that at each stage of its development from egg to adult an organism must be an integrated, functioning whole. Another is that for any form to arise in an organism at a time, it must develop from the materials and processes at the organism’s disposal. The requirement of integration and the processes of development that produce it leave their distinctive marks on form. It seems reasonable, then, to suppose that one might appeal to the processes of development and the principles by which they operate in explaining the nature and distribution of biological form. (See here Gould 1977; Rose and Lauder 1996) John Dupre', Comments on Mitchell and Arnold Jonathon Hodge, Comments on Lennox and Love These issues (and more) are raised in Gould and Lewontin (1979).