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Chapter 54 – Ecosystems TJ Biology Olympiad Cassandra Xia Trophic structure – feeding systems Divided into trophic levels- based on main source of nutrition Primary Producers—autotrophs - photoautotrophs plants phytoplankton (algae and bacteria) multicellular algae and aquatic plants - chemoautotrophic bacteria which oxidizes H2S Autotrophs- need only CO2 as a carbon source Heterotrophs- needs at least one organic nutrient as a source of carbon for making other organic compounds Primary Consumers—herbivores - eat plants or algae - Opportunistic – supplement diet with heterotrophic material if available (e.g. with eggs and hatchlings) Secondary consumers - carnivores that eat herbivores Tertiary consumers - carnivores that eat carnivores Photo- derives energy from light Chemo- derives energy from chemicals taken from the environment (animals, fungi, prokaryotes, and protists are chemoheterotrophs) Detritivores - eat detritus (non-living organic material = feces, fallen leaves, dead organisms) Gross primary productivity- the amount of light energy converted to chemical energy (organic compounds) by autotrophs of an ecosystem during a time period; the rate at which light energy is converted to chemical energy in an ecosystem - not all stored as organic material because some used in cell respiration Standing crop biomass- total - measured in g/m2/yr of dry weight or J/ m2/yr biomass of photoautotrophs Net primary productivity- what is stored as organic material present at a given time. Measured - NPP = GPP – Respiration in grams/kilograms. - Available to consumers in the ecosystem - NPP:GPP ratio smaller for large producers which have to support large stem and root systems More on primary productivity - Open ocean has low NPP by area, but contributes a lot to Earth’s NPP due to huge surface area - Estuaries and reefs have high NPP by area, but contributes little to Earth’s NPP due to small suface area - Tropical rain forest have high NPP by area, and contributes a lot to Earth’s NPP yay - Limiting nutrient- the thing that must be added in order for production to increase in a particular area usually nitrogen or phosphorus sometimes light Secondary Productivity- the rate at which an ecosystem’s consumers convert the chemical energy of the food they eat into their own new biomass - Energy in a consumer is lost in: o Cell respiration - o Feces o Growth (available energy for the next level) Carnivores are slightly more efficient at converting energy into biomass because meat is easier to digest. BUT animals unlike plants use up a lot of energy running around and stuff, so if all humans were to go vegetarian there would be enough food for everyone to eat Ecological efficiency- percentage of energy transferred from one trophic level to the next (varies from 5% to 20% but use the “10% rule”) Ecological pyramids - Mostly all bottom-heavy - Types: o Productivity pyramid o Pyramid of numbers o Biomass pyramid Turnover time = Standing crop biomass (kg/m2) / Productivity (kg/m2/day) Water Cycle - Evaporation over the ocean, evapotranspiration over land - Clouds rain over ocean and land - Runoff and groundwater return to ocean (percolation through soil) Carbon Cycle - Photosynthesis and respiration have opposing effects on CO2 levels - Because of the burning of locked away carbon in the form of fossil fuels, CO2 levels steadily increasing and contributing to the Greenhouse Effect - CO2 levels have seasonal fluctuation: o Lowest during summer o Highest during winter o Why? - In the ocean, it is more complicated: o The ocean has reactions between limestone (CaCO3), water, and CO2 o H2O + CO2 <=> H2CO3 H2CO3 + CaCO3 <=> Ca(HCO3)2 <=> Ca2+ + 2HCO3+ 22HCO3 <=> 2H + 2CO3 o As CO2 is used for photosynthesis in marine environments, the first equation shifts to the left. o Some aquatic autotrophs can use HCO3o The amount of carbon in the ocean is 50 times that in the atmosphere, the ocean may be an important carbon sink as carbon dioxide levels increase Phosphorous Cycle - Organisms need phosphorus for o Nucleic acids o Phospholipids o ATP o Bones & teeth - Weathering of rocks adds phosphate to the soil - Plants absorb phosphorus in the inorganic form of phosphate (PO43-) Consumers get it by eating plants Decomposers return it to the soil Also, phosphorus leaches into the water table, draining into the sea There it becomes incorporated into sedimentary rocks that may be part of a terrestrial ecosystem later in time Phosphates in the form of sewage and runoff fertilizer causes eutrophication in aquatic systems Nitrogen Cycle- this is probably the most quizzed thing on the whole exam. A question on the nitrogen cycle is practically guaranteed. It is worth learning. - Nitrogen is needed to build amino acids and therefore proteins. - Nitrogen is the most abundant element in the Earth’s atmosphere, but it cannot be used by most organisms if it is in the form N2 - N2 is converted into more usable forms in three main ways o By nitrogen fixation- free-living soil bacteria, cyanobateria, and bacteria in root nodules of legumes converts N2 to NH3 o Atmospheric deposition (lightning)converts to NH4+ or NO3- which dissolves in rain and gets added to the soil o Industrial production of fertilizer - Most soil is slightly acidic so NH3 produced by nitrogen fixation picks up an H+ and becomes NH4+ - Plants can absorb NH4+ but they can also absorb NO3- Nitrification by nitrifying bacteria converts NH4+ to NO3- Animals get their nitrogen by eating plants - Cycle part: o Denitrification- denitrifying bacteria convert NO3- back to N2 o Ammonification- bacterial and fungal decomposers convert organic nitrogen in plants and animals to NH4+ Decomposition rates depend on: Temperature Availability of H2O and O2 Soil chemistry Fires Fast Slow Long Term Ecological Research (LTER) at Hubbard Brook - Forest logged and herbicide sprayed - Water runoff increased - Ca2+ x 4 loss, K+ x 15 loss, NO3- x 60 loss Tropical rain forests Temperature forests Tundra months-years 4-6-years 50 years Environmental Hazards Agriculture and Nitrogen Cycling - Move nutrients from one area to another - Nutrients lost through shipping agriculture and erosion - Fertilizers must be added for “free period” (no need to add nutrients) Toxins - Many cannot be degraded by microorganisms - Some compounds may be converted to toxins CH3—Hg e.g. Hg bacteria - insoluble soluble, accumulates in tissues of fish and humans Chlorinated hydrocarbons (DDT and PCBs) are indicated in endrocrine disruption Undergo biological magnification- increased concentrations of toxins in successive trophic levels of a food web caused by eating toxins concentrated in the tissues of lower level organisms DDT and DDE (product of partial breakdown) interfere with calcium deposition in egg shells of eagles, pelicans, and ospreys Accelerated Eutrophication of Lakes - Cultural eutrophication is eutrophication caused by humans. Sources of it include Sewage Factory wastes Runoff of animal waste Fertilizers - Algae blooms - Some algae die - Decomposing bacteria use up O2 Carbon Dioxide and the Greenhouse Effect - Greenhouse effect: gases absorb reradiate infrared radiation back at the earth, increasing temperature (without it, earth’s surface temperature would be -18° C) - More CO2 is being released by burning fossil fuels and is contributing to global warming o Melt polar ice o Alter precipitation patterns (central US drier) - With more CO2, plants become more productive but the higher levels favor C3 plants (like wheat + soy) over C4 plants (like corn) causing C3 plants to spread into areas originally covered by C4 plants Depletion of Atmospheric Ozone - Chlorofluorocarbons (refrigerants) is a catalyst in the reaction O3 → O2 - Cold temperature over Antarctica facilitate reactions