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Ecology Patterns in the Environment • Where an organism lives is called it’s Habitat. • It is shaped by the organisms interaction with abiotic and biotic factors. habitats Niche • An organisms niche is its functional position in the environment. It includes a description of how the organism interacts with the abiotic and biotic elements of the environment. Niche Law of Tolerance- abiotic factors The law of tolerance states that “For each abiotic factor, an organism has a range of tolerances within which it can survive.” Tolerance range Number of organisms Optimum range Unavailable niche Marginal niche Examples of abiotic factors that influence size of the realized niche: Preferred niche Marginal niche Unavailable niche Biotic factors -Competition and Niche Size Competition affects the size of a competitor’s realized niche. The effect is dependent on the intensity and type of the competition. Niches are narrower with moderate interspecific competition (Fig. 1). Realized niche of species Fig. 1 Narrower niche Fig. 2 Intense interspecific competition results in a very narrow realized niche as species specialize to exploit a narrower range of resources (Fig. 2). Intense intraspecific competition results in a broader realized niche as individuals are forced to occupy suboptimal conditions (Fig. 3). Broader niche Fig. 3 Possible tolerance range Gause’s Principle Gause’s competitive exclusion principle states: “two or more resource-limited species, having identical patterns of resource use, cannot coexist in a stable environment: one species will be better adapted and will outcompete or otherwise eliminate the other(s)”. Amount eaten If two species compete for some of the same resources (e.g. food items of a particular size), their resource use curves will overlap. In the zone of overlap, interspecific competition is the most intense. Zone of overlap Species A Species B Resource use as measured by food item size Competing species cannot occupy the same niche Barnacles • Chthamalus and Balanus compete for space. • Note that Chthamalus realised niche is much smaller than its fundamental niche as it is out competed by Balanus. Niche Differentiation Interspecific competition is usually less intense than intraspecific competition because niche overlap between species is not complete. Species with similar ecological requirements may reduce competition by exploiting different microhabitats within the ecosystem. Example: Ecologically similar damsel fish at Heron Island, Queensland, Australia exploit different resources or regions over the coral reef. Pw Pomacentrus wardi Pf Pomacentrus flavicauda Pb Pomacentrus bankanensis Sa Stegastes apicalis Pl Plectroglyphidodon lacrymatus Ef Eupomacentrus fasciolatus Eg Eupomacentrus gascoynei Gb Glyphidodontops biocellatus Reef crest Sea level Zonation in a lake Zonation- changing environments are often divided into zones Stratification in a forest is a type of vertical zonation Different organisms will occupy different niches within the different zones Succession • Communities change over time. • Early communities alter the environment allowing other organisms to colonise. • Eventually a climax community will form Secondary succession occurs after a catastrophe or something that disrupts the environment Succession and stratification • http://www.bbc.co.uk/ nature/habitats/Tropica l_and_subtropical_mois t_broadleaf_forests#p0 0fdh3v • http://www.bbc.co.uk/ nature/habitats/Tropica l_and_subtropical_conif erous_forests#p00ffnqd • https://www.youtube.c om/watch?v=k03vxRYsJ 4Y • https://www.youtube.c om/watch?v=CQ2Xl6Zq zRI • https://www.youtube.c om/watch?v=Ax6zlSzyNM