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Transcript
Chapter 45: Sensory Systems
CHAPTER 45
Sensory Systems
Chapter 45: Sensory Systems
Chapter 45: Sensory Systems
Sensory Cells, Sensory Organs, and
Transduction
Chemoreceptors: Responding to Specific
Molecules
Mechanoreceptors: Detecting Stimuli that
Distort Membranes
Chapter 45: Sensory Systems
Chapter 45: Sensory Systems
Photoreceptors and Visual Systems:
Responding to Light
Sensory Worlds Beyond Our Experience
Chapter 45: Sensory Systems
Sensory Cells, Sensory
Organs, and Transduction
• Sensory cells transduce information about
an animal’s external and internal
environment into action potentials.
Review Figures 45.1, 45.2
4
Chapter 45: Sensory Systems
figure 45-01a.jpg
Figure
45.1 – Part
1
Figure 45.1 – Part 1
Chapter 45: Sensory Systems
figure 45-01b.jpg
Figure 45.1
– Part 2
Figure 45.1 – Part 2
Chapter 45: Sensory Systems
figure 45-02.jpg
Figure
45.2
Figure 45.2
Chapter 45: Sensory Systems
Sensory Cells, Sensory
Organs, and Transduction
• The interpretation of action potentials as
particular sensations depends on which
neurons in the CNS receive them.
8
Chapter 45: Sensory Systems
Sensory Cells, Sensory
Organs, and Transduction
• Membrane receptor proteins of sensory cells
cause ion channels to open or close,
generating receptor potentials.
• Receptor potentials can spread to regions of
the sensory cell plasma membrane that
generate action potentials, or influence
release of neurotransmitter from the
sensory cell.
Review Figure 45.3
Chapter 45: Sensory Systems
figure 45-03.jpg
Figure
45.3
Figure 45.3
Chapter 45: Sensory Systems
Sensory Cells, Sensory
Organs, and Transduction
• Adaptation enables the nervous system to
ignore irrelevant stimuli while remaining
responsive to relevant or to new stimuli.
11
Chapter 45: Sensory Systems
Chemoreceptors: Responding
to Specific Molecules
• Smell, taste, and the sensing of pheromones
are examples of chemosensation.
• Chemoreceptor cells have receptor proteins
that can bind to specific molecules that
come into contact with the sensory cell
membrane.
Review Figures 45.5, 45.6
12
Chapter 45: Sensory Systems
figure 45-05.jpg
Figure
45.5
Figure 45.5
Chapter 45: Sensory Systems
Chemoreceptors: Responding
to Specific Molecules
• Binding of an odorant molecule to a
receptor protein causes production of a
second messenger in the chemoreceptor
cell.
• The second messenger alters ion channels
and creates a receptor potential.
15
Chapter 45: Sensory Systems
Chemoreceptors: Responding
to Specific Molecules
• Chemoreceptors in the mouth cavities of
vertebrates are responsible for the sense of
taste.
Review Figure 45.6
16
Chapter 45: Sensory Systems
figure 45-06.jpg
Figure
45.6
Figure 45.6
Chapter 45: Sensory Systems
Mechanoreceptors: Detecting
Stimuli that Distort Membranes
• The skin has a diversity of
mechanoreceptors that respond to touch
and pressure.
• The density of mechanoreceptors in any
skin area determines the sensitivity of that
area.
Review Figure 45.7
18
Chapter 45: Sensory Systems
figure 45-07.jpg
Figure 45.7
Figure 45.7
Chapter 45: Sensory Systems
Mechanoreceptors: Detecting
Stimuli that Distort Membranes
• Stretch receptors in muscles, tendons, and
ligaments inform the CNS of the positions of
and the loads on parts of the body.
Review Figure 45.8
20
Chapter 45: Sensory Systems
figure 45-08.jpg
Figure
45.8
Figure 45.8
Chapter 45: Sensory Systems
Mechanoreceptors: Detecting
Stimuli that Distort Membranes
• Hair cells are mechanoreceptors that are not
neurons.
• Bending of their stereocilia alters their
membrane proteins and therefore their
receptor potentials.
• Hair cells are found in organs of equilibrium
and orientation.
Review Figures 45.9, 45.10, 45.11
22
Chapter 45: Sensory Systems
figure 45-09.jpg
Figure
45.9
Figure 45.9
Chapter 45: Sensory Systems
figure 45-10.jpg
Figure
45.10
Figure 45.10
Chapter 45: Sensory Systems
figure 45-11a.jpg
Figure
45.11 – Part
1
Figure 45.11 – Part 1
Chapter 45: Sensory Systems
figure 45-11b.jpg
Figure 45.11
– Part 2
Figure 45.11 – Part 2
Chapter 45: Sensory Systems
Mechanoreceptors: Detecting
Stimuli that Distort Membranes
• Hair cells are responsible for mammalian
auditory sensitivity.
• Ear pinnae collect and direct sound waves
to the tympanic membrane, which vibrates
in response to sound waves.
• The vibrations are amplified through a chain
of ossicles that conduct them to the oval
window.
• Movements of the oval window create
pressure waves in the fluid-filled cochlea.
Review Figure 45.12, 45.13
29
Chapter 45: Sensory Systems
figure 45-12a.jpg
Figure 45.12
– Part 1
Figure 45.12 – Part 1
Chapter 45: Sensory Systems
figure 45-12b.jpg
Figure 45.12
– Part 2
Figure 45.12 – Part 2
Chapter 45: Sensory Systems
figure 45-12c.jpg
Figure
45.12 – Part
3
Figure 45.12 – Part 3
Chapter 45: Sensory Systems
Mechanoreceptors: Detecting
Stimuli that Distort Membranes
• The basilar membrane running down the center of
the cochlea is distorted at specific locations
depending on the frequency of the pressure wave.
• These distortions cause bending of hair cells in the
organ of Corti, which rests on the basilar
membrane.
• Changes in hair cell receptor potentials create
action potentials in the auditory nerve, which
conducts the information to the CNS.
Review Figure 45.13
31
Chapter 45: Sensory Systems
figure 45-13.jpg
Figure 45.13
Figure 45.13
Chapter 45: Sensory Systems
Photoreceptors and Visual
Systems: Responding to Light
• Photosensitivity depends on the capture of
photons of light by rhodopsin.
• Rhodopsin is a photoreceptor molecule
consisting of opsin and retinal.
• Absorption of light by retinal is the first step
in a cascade of intracellular events leading
to a change in the receptor potential of the
photoreceptor cell.
Review Figure 45.15
33
Chapter 45: Sensory Systems
figure 45-14.jpg
Figure
45.14
Figure 45.14
Chapter 45: Sensory Systems
figure 45-15.jpg
Figure
45.15
Figure 45.15
Chapter 45: Sensory Systems
Photoreceptors and Visual
Systems: Responding to Light
• When excited by light, vertebrate
photoreceptor cells hyperpolarize and
release less neurotransmitter onto neurons
with which they form synapses.
• They do not fire action potentials.
Review Figures 45.16, 45.17, 45.18
35
Chapter 45: Sensory Systems
figure 45-16.jpg
Figure
45.16
Figure 45.16
Chapter 45: Sensory Systems
figure 45-17a.jpg
Figure 45.17
– Part 1
Figure 45.17 – Part 1
Chapter 45: Sensory Systems
figure 45-17b.jpg
Figure
45.17 –
Part 2
Figure 45.17 – Part 2
Chapter 45: Sensory Systems
Photoreceptors and Visual
Systems: Responding to Light
• Vision results when eyes focus patterns of
light onto layers of photoreceptors.
• The simple eye cups of flatworms can sense
the direction of a light source
• The compound eyes of arthropods can
detect shapes and patterns
• The eyes of cephalopods and vertebrates
have lenses.
Review Figures 45.18, 45.19, 45.20
40
Chapter 45: Sensory Systems
figure 45-18.jpg
Figure
45.18
Figure 45.18
Chapter 45: Sensory Systems
figure 45-19.jpg
Figure 45.19
Figure 45.19
Chapter 45: Sensory Systems
figure 45-20.jpg
Figure 45.20
Figure 45.20
Chapter 45: Sensory Systems
Photoreceptors and Visual
Systems: Responding to Light
• The eyes of vertebrates and cephalopods
focus detailed images of the visual field onto
dense arrays of photoreceptors that
transduce the visual image into neural
signals.
Review Figure 45.21
43
Chapter 45: Sensory Systems
figure 45-21.jpg
Figure
45.21
Figure 45.21
Chapter 45: Sensory Systems
Photoreceptors and Visual
Systems: Responding to Light
• The vertebrate photoreceptors are rod and
cone cells.
• Rod cells are responsible for dim light and
black-and-white vision
• Cone cells are responsible for color vision by
virtue of their spectral sensitivities.
Review Figure 45.23
45
Chapter 45: Sensory Systems
Photoreceptors and Visual
Systems: Responding to Light
• The vertebrate retina is a dense array of neurons
lining the back of the eyeball.
• It consists of five layers of cells.
• The outermost layer consists of rods and cones.
• The innermost layer consists of ganglion cells,
which send their axons in the optic nerve to the
brain.
• Between the photoreceptors and ganglion cells are
neurons that process information from the
photoreceptors.
Review Figure 45.24
47
Chapter 45: Sensory Systems
figure 45-24.jpg
Figure
45.24
Figure 45.24
Chapter 45: Sensory Systems
Photoreceptors and Visual
Systems: Responding to Light
• The area of the retina that receives light
from the center of the visual field, the
fovea, has the greatest density of
photoreceptors.
• In humans it contains almost exclusively
cone cells, which are responsible for color
vision but not very sensitive in dim light.
50
Chapter 45: Sensory Systems
Photoreceptors and Visual
Systems: Responding to Light
• Each ganglion cell is stimulated by light
falling on a small circular patch of
photoreceptors called a receptive field.
• Receptive fields have a center and a
surround, which have opposing effects on
the ganglion cell.
• If the center is excitatory, the surround is
inhibitory, and vice versa.
See fig.45.25
51
Chapter 45: Sensory Systems
figure 45-25a.jpg
Figure
45.25 –
Part 1
Figure 45.25 – Part 1
Chapter 45: Sensory Systems
figure 45-25b.jpg
Figure
45.25 –
Part 2
Figure 45.25 – Part 2
Chapter 45: Sensory Systems
Sensory Worlds Beyond Our
Experience
• Many animals have sensory abilities that we
do not share.
• Bats echolocate
• Insects see ultraviolet radiation
• Pit vipers “see” infrared radiation
• Fish sense electric fields.
52