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Zoology Unit 2 Evolution – the change in a population over time. Pre-Darwin Ideas 1. Lamarck A. French zoologist Thought organisms developed new organs or modified existing organs to adapt to new situations. Organs not used would degenerate. Species did not become extinct, only evolved into new species. Giraffes – Thought they got long necks by stretching to get to higher leaves. Wrong because modifications like that cannot be passed to the next generation. 2. Darwin’s Ideas A. Thought species changed over long periods of time. (too long) B. Theory of Uniformitarianism (Charles Lyell) said the earth was very old and the earth changed over time. This idea reinforced Darwin’s idea. C. From looking at fossils on South America he recognized that some animals become extinct without leaving descendents or becoming new species. D. On Galapagos he observed finches and tortoises. E. The finches have different size beaks with different shapes. Depending on what they eat. They show “adaptive radiation”. Adaptive radiation is the formation of new forms of a species from an ancestral species in response to the opening of new habitats. F. Natural selection – developed after reading Thomas Malthus. He said human population will continue to grow until stopped by some factor such as war, famine, disease, etc. Darwin thought the same thing might happen in nature. G. Natural Selection has 4 parts. 1. All organisms produce more offspring than can reproduce. 2. Inherited variation exists. Variations come from mutations in DNA and random mating. Some may be advantageous, some may be harmful. 3. Individuals with the advantageous traits are more likely to survive and pass them to the next generation. These are called adaptive traits. 4. Adaptive traits are passed on and become more frequently seen in the population. Traits that are dis-advantageous to the individual will eventually be eliminated. 3. MICRO and MACRO EVOLUTION A. Microevolution is a change in the frequency of alleles in a population over time. Examples are drug resistant strains of bacteria and bug that are resistant to pesticides. B. Macroevolution is the large scale changes that result in extinction and the formation of new species. C. What scientists refer to as evidence for macroevolution. 1. Biogeography is the study of the geographic distribution of plants and animals. Biogeographers try to explain: Why similar animals end up in places that are not connected physically. Ex. – cats Why animals separated by geographical barriers are often very different in spite of similar environmental conditions. Ex – Australia and New Zealand. 2. Paleontology – the study of fossil records. Older fossils are usually found below or deeper than new fossils. By looking at how fossils change from older layers to newer layers we see how animals have changed. 3. Comparative Anatomy Convergent evolution occurs when we find superficially similar structures in unrelated organisms. Ex – wings in birds and insects. In this case the wings are said to be analogous structures. Comparative anatomy is the study of living and fossilized animals and their relationships. In comparative anatomy homologous structures are studied. These are structures in different animals that suggests they have a common ancestor. Ex. bones in the forelimbs of birds, humans, lizards, bats, etc. as seen in figure 4.11 on page 62. Homologous DNA molecules – related animals have similar DNA derived from the ancestral DNA. 4. Developmental Patterns By looking at early embryonic stages we can see similarities. From the embryonic stage organisms begin to develop differently. This is due to genes that control how fast the bones and organs develop and how they are proportioned. Phylogeny – Scientists use evolutionary evidence to build a phylogentic tree. Phylogeny refers to the evolutionary relationships of modern descendents of a common ancestor. The tree shows the relationship between organisms, the branches show where species diverged into separate species. A. Population 6+ and Gene Pool 1. A population is a group of individuals of the SAME species that occupy an area at the same time. Within the population the individuals will all have the same kind of genes. Differences seen in individuals is due to the presence of different alleles. All the alleles for traits in all traits in a population is called the gene pool. It is a pool of hereditary resources containing all the possible alleles an individual can be made from. 2. Population genetics is the study of the genetic events in the gene pool. The HardyWeinberg theorem describes what happens to the frequency of alleles in a population over time. The Hardy-Weinberg theorem says if certain things (assumptions) are met no evolution will take place and the frequency of alleles will not change in future generations. What are these assumptions that must be met to prevent evolution? There are four. First – The population size must be large. Second – Mating between individuals must be random. Natural selection must take place. Third – Individuals cannot move into or out of the population. This keeps new alleles from entering the gene pool. Fourth – No new types of alleles are introduced due to mutation. It is very unusual for all these assumptions to be met so most populations are evolving. B. Evolutionary Mechanisms Things that can happen when the Hardy-Weinberg assumptions are not met. 1. Sometimes individuals meet and reproduce by chance. No natural selection is involved. When this changes the frequency of alleles in a population it is called “genetic drift” . 2. Genetic drift is most likely to occur in small populations. 3. The Founder Effect is an example of genetic drift. This occurs when a very few individuals from a large population colonize a new environment away from the parent population. The new colony will only contain the genetic make-up of the individuals that founded the new colony. 4. Another type of genetic drift is the Bottleneck Effect. This occurs when an event drastically reduces the size of a population. The remaining population may grow to its original size but it will not have the same genetic diversity. C O G Z I Y N K P M M T K N H I O Z R G C C C R H T Y D P Y G R H N Z G P R G H P H G R P N P N P R H H Z G H Z G N N N H Z R G Z Z P Z P Most biologist believe genetic diversity is important to a species survivability. This is because a population with high genetic diversity will allow it to survive environmental changes. 5. Gene Flow – The HardyWeinberg theorem assumes no immigration or emigration. If it does occur there will be a change in the number of alleles and the kinds of alleles in a population. 6. Mutation – mutations are the origin of new alleles and variations. These may help the organism become better adapted to their environment. Or not. C. SPECIES AND SPECIATION 1. A species is a group of populations in which genes are actually or potentially exchanged through successful interbreeding. 2. Speciation is the formation of new species. This can occur in different ways such a reproductive isolation. C. Rates of Evolution 1. Phylogentic gradualism – in this model changes are gradual over long periods of time. 2. Punctuated equilibrium – in this model there are long periods of stasis then rapid periods of change. The sudden and rapid change may be due to environmental changes (like climate change).