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Classification & Evolution Evolution - Microevolution A change within the gene pool of a population over time (changes in the percentages of alleles) Lamarck vs. Darwin The Theory of Evolution Darwin Descent with Modification: Darwin - Organisms can have offspring that are modified forms of themselves. Modern Interpretation – DNA mutations and genetic recombination through meiosis and fertilization cause variations within populations Natural Selection Organisms have more babies than can be supported by the environment The most successful at surviving and reproducing pass those genes on and become more numerous in the population Therefore, evolution is due to multiple forms within populations – if an uncommon form works better – it will become more common over time (populations change over time to what works the best) These smaller changes become bigger over time. The commonalities among organisms are evidence of common ancestry. Selection Factors (Pressures) Predator/Prey – ability to get prey or evade predators Competition – better competitor for food, homes, mates Abiotic Factors (amt. of ppt., wind, type of soil, amt. of sunlight, temperature) Sex Selection The five mechanisms of evolution – It’s not just Natural Selection 5 Fingers of Evolution - TedEd Small population (Genetic Drift) – next generation doesn’t match probability so get a change in the allelic frequencies due to chance Non-random mating – type of selection Mutation Gene Flow – things moving in or out of a population changing the allelic frequencies Natural Selection – most important and only one that makes improvements! Examples of the 5 mechanisms of Evolution Natural Selection is the only mechanism of evolution that creates Adaptations!! Adaptations: Genetic traits that allow organisms to survive better in their environment Even though natural selection causes things to generally improve and become more adapted to their environment, evolution doesn’t always create the “perfect” best adapted forms if the “perfect” random mutation doesn’t happen Why biodiversity remains Even though natural selection tends to eliminate diversity for the best suited forms – diversity is maintained by: Meiosis and genetic recombination Heterozygosity Continued mutations Neutral mutations Multiple Selection Factors Continuously changing environments This is important because if environment changes – there will be a part of the population or species that can survive Evidence of Evolution 1. Direct evidence: Finches Fish peppered moths Insects bacteria Evidence of Evolution Continued 2. Artificial Selection Pigeons Mustard Plant Fossils – see new things and see many things disappear Fossil Record – fossils show descendancy – relatedness matches age of fossils 3. Don’t find different vertebrate classes in the same age rock – appears to happen chronologically Can find transitional fossils linking ancient and modern species See fossils of things that no longer exist and things that weren’t in the older fossils – appears to be new Evidence of Evolution Continued 4. Taxonomy (The Study of Classification) Shows unity and diversity – everything seems to be related and modified versions of each other Taxonomy – the study of classification Why do we classify things? Helps us understand how things are related which helps us understand things in general Linnaeus came up with a classification system to better understand living things The system ended up being a hierarchy What is a hierarchy? Classify shoes with a partner Is this a hierarchy? What does it help us understand about shoes? What is the classification scheme for living things like? The classification system Domain, Kingdom, Phylum, Class, Order, Family, Genus, Species (Kings play cards on fine green sheets) The hierarchy shows a unity that all living things are related and share a common ancestor (seem to be modified versions of each other) It also shows diversity – that they have changed a great deal over time Biological Taxa Example of Classification Goes from general to specific Continually branches The longer things stay together before branching, the more they have in common If two things are in the same category – they share everything more general in common Taxonomy All living things have some things in common Some have more things in common and more closely related The further down the scheme they stay together (from general to specific), the more traits they share in common and the more related they are How do we Classify to show evolutionary relationships? Comparative Anatomy Homologous Structures – structurally similar even though have different functions (shows relatedness vs. individual engineering) Vestigial Organs – “left-overs” – no funtion in current times Comparative Embryology – the longer in development things look the same the more related – ex. all vertebrates go through the same stages early on – changes in genes and reading of genes causes different forms Biochemistry/Molecular Biology – same DNA in all organisms – looks like modified copies of each other (mutations to make different proteins) Comparative Anatomy Analogous structures – similar due to like environments, built from different structures (ex. Wings or birds and insects) DO NOT SHOW RELATEDNESS – NOT USED FOR CLASSIFICATION Homologous structures – similar due to common structure and therefore common ancestry (ex. Wing of bat, whale fin, arm of human, paw of dog) Use ONLY homologous structures for classification Problem with comparative anatomy – like structures not necessarily from common ancestor – may be shaped by same environmental factors Homologous Structures Homologous Structures show evolutionary relationships and should be used for classification Analogous structures do not show evolutionary relationship and are not used for classification Comparative Embryology Living things share some common embryological development – the longer the same, the more related Comparative Biochemistry All things show some DNA and protein relatedness – the more DNA and proteins they have in common – the more closer related. If they have the same protein, more amino acids in common in the protein – the more related Comparative Biochem If have same proteins, closer their DNA sequences – the more alike Dolphins are closer related to bats than sharks – can tell through proteins and DNA that they are more related Evolution – a step further Microevolution – change in the gene pool within a population (change in allelic frequencies) Speciation – creation of new species Macroevolution – creating all of the various taxa from a common ancester Species Concept Species – organisms that interbreed and produce fertile offspring under natural conditions Speciation – the formation of new species Species concept - Why species remain distinct: Habitat Isolation – live in different areas Behavioral Isolation – mating rituals, firefly lighting patterns Temporal Isolation – different mating times (seasonal), different times of flowering, nocturnal vs. day Mechanical Isolation – physically impossible to mate Gametic Isolation – gametes can’t match up Infertility of Hybrids Speciation – see the appearance of new species in fossils – where did they come from? Production of new species due to separation migration to different islands new mountain separates them lake dries up to multiple little ponds Separated groups undergo different mutations Each separated part of the species is acted upon by different “selectors” and eventually become so different that they can’t or do not any longer interbreed Examples of Speciation Why evolution takes places once a population becomes separated Organisms on the edge are usually different anyway Small group leaving may not be representative of whole More change due to chance in small groups Neutral mutations may become fixed without selective pressures due to small population size Different mutations Different selective pressures Therefore: Microevolution over time slowly changes each population until they are different species Macroevolution substantial change in organisms Origin of new phyla, classes, orders, families Is it due to the cumulative product of microevolution or some big event or…???? The appearance of flowering plants seems to be all at once????? The appearance of mammals seems to be all at once????? Punctuated Equilibrium Big changes (episodes of speciation) followed by slow gradual change (if optimized for environment – shouldn’t be a lot of change due to selection unless large change in selection pressures) Due to quick geographic separation and genetic drift Due to sudden genome changes Changes may not be shown in fossils Mechanisms of Macroevolution Pre-adaptation – structure is adapted for 1 thing and later used for another function (gradual change in existing structure leads suddently to a new function) Example – lattice-like bones of birds – some dinosaurs had it but must have had another function Changes in developmental genes Mass Extinction – due to huge geographical changes (climate, destruction of habitats) – leaves it open for species to fill new places Mechanisms of Macroevolution Cont. Accumulation of Microevolution not preserved in fossil record or intermediates not found due to small numbers Summary of Macroevolution May be due to rapid changes: Mass separations Rapidly changing environments Chromosomal or developing genes mutating Mutations acted upon by huge genetic drift and selection Mass extinctions causing adaptive radations Remaining Questions about Macroevolution Could it really be compounded microevolution? What is gradual vs. quick? What is missing from the fossil record? Do different mechanisms work at different levels?