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Ecosystems An ecosystem is all of the organisms in an area, along with their nonliving environment Example: aquarium Living + Non-living (Biotic + Abiotic) Trophic Levels Organisms in a community are related to each other through feeding relationships Each step up in the transfer of energy is known as a trophic level All energy ultimately comes from the SUN Trophic Levels Decomposers/ Detritivores Eat detritus (organic waste/remains of dead organisms) Can fit in to a food chain or web at any location Trophic Levels Producers Convert solar (or chemical) energy into organic compounds Primary consumers Eat producers Secondary consumers Eat primary consumers Tertiary consumers Eat secondary consumers Pyramid of Numbers/Biomass/Ener gy Numbers, energy, & biomass decreases as one moves up the food chain. Biomass- dry mass of organic matter Trophic Levels Ten-Percent Law Usable energy is lost through each transfer of energy Why? (Remember the law of conservation of energy says energy cannot be created or destroyed; it only changes form.) Only about 10% of the energy at one trophic level is transferred to the next trophic level. 90% is lost as heat with each transfer. Food Chain A straight-line marsh hawk upland sandpiper sequence of who eats whom garter snake Simple food chains are rare in nature cutworm plants Tall-Grass Prairie Food Web marsh hawk sandpiper crow snake frog weasel badger coyote spider sparrow earthworms, insects vole pocket gopher grasses, composites ground squirrel Primary Productivity Primary Productivity: The amount of light energy converted to sugars by autotrophs in an ecosystem Gross vs. Net Primary Productivity GPP: the amount of light energy that is converted to chemical energy by photosynthesis per unit time NPP: GPP minus the energy used by the primary producers for cellular respiration GPP-R=NPP Limiting Nutrients What limits primary production? Aquatic Ecosystems Light (depth penetration) Nitrogen Phosphorus Terrestrial Ecosystems Temperature Moisture Minerals (N & P are the main limiting factors for plants.) Biogeochemical Cycle The flow of a nutrient from the environment to living organisms and back to the environment Main reservoir for the nutrient is in the environment Hydrologic Cycle Atmosphere wind-driven water vapor 40,000 evaporation precipitation from ocean into ocean 425,000 385,000 precipitation onto land 111,000 evaporation from land plants (evapotranspiration) 71,000 surface and groundwater flow 40,000 Ocean Land Figure 48.14 Page 876 diffusion between atmosphere and ocean bicarbonate and carbonate in ocean water photosynthesis combustion of fossil fuels aerobic respiration marine food webs death, incorporation sedimentation into sediments uplifting sedimentation marine sediments Carbon Cycle - Marine Figure 48.16 Page 878 atmosphere combustion of fossil fuels volcanic action terrestrial rocks weathering photosynthesis aerobic combustion respiration of wood sedimentation land food webs soil water leaching, runoff death, burial, compaction over geologic time Carbon Cycle - Land peat, fossil fuels Figure 48.16 Page 878 Carbon in Atmosphere Atmospheric carbon is mainly carbon dioxide Carbon dioxide is added to atmosphere Aerobic respiration, volcanic action, burning fossil fuels Removed by photosynthesis Greenhouse Effect Greenhouse gases impede the escape of heat from Earth’s surface Figure 48.18, Page 880 Global Warming Long-term increase in the temperature of Earth’s lower atmosphere Figure 48.19, Page 881 Nitrogen Cycle Nitrogen is used in amino acids and nucleic acids Main reservoir is nitrogen gas in the atmosphere Nitrogen Cycle gaseous nitrogen (N2) in atmosphere nitrogen fixation by industry food webs on land uptake by excretion, death, uptake by fertilizers autotrophs decomposition autotrophs nitrogen fixation NH3-,NH4+ in soil leaching nitrogenous wastes, remains NO3in soil dentrification ammonification 2. Nitrification 1. Nitrification NO2in soil leaching Figure 48.21 Page 882 Nitrogen Fixation Plants cannot use nitrogen gas Nitrogen-fixing bacteria convert nitrogen gas into ammonia (NH3) Ammonia and ammonium can be taken up by plants Ammonification & Nitrification Bacteria and fungi carry out ammonification conversion of nitrogenous wastes to ammonia Nitrifying bacteria convert ammonium to nitrites and nitrates Nitrogen Loss Nitrogen is often a limiting factor in ecosystems Nitrogen is lost from soils via leaching and runoff Denitrifying bacteria convert nitrates and nitrites to nitrogen gas Phosphorus Cycle Phosphorus is part of phospholipids and all nucleotides It is the most prevalent limiting factor in ecosystems Main reservoir is Earth’s crust; no gaseous phase Phosphorus Cycle mining FERTILIZER GUANO excretion agriculture uptake by autotrophs MARINE FOOD WEBS weathering DISSOLVED IN OCEAN WATER uptake by autotrophs weathering DISSOLVED IN SOILWATER, LAKES, RIVERS death, decomposition sedimentation LAND FOOD WEBS death, decomposition settling out leaching, runoff uplifting MARINE SEDIMENTS TERRESTRIAL ROCKS over geologic time Figure 48.23, Page 884 Human Impact on Ecosystems Increased Eutrophication of Lakes Increase in nutrient levels (phosphates, nitrates, etc.) Can lead to algal blooms Hypoxia What is it? Why? Can lead to the eventual loss of fish and other aquatic organisms Accelerated by sewage/factory wastes, leaching of fertilizers into freshwater Human Impact on Ecosystems Combustion of Fossil Fuels Leads to acid precipitation Changes the pH of aquatic ecosystems and affects the soil chemistry of terrestrial ecosystems Human Impact on Ecosystems Biological Magnification Toxins become more concentrated as they move up the food chain Toxins that are lipophilic cannot be excreted in urine (water!), so they are stored in fatty tissue (adipose tissue) unless the organism has enzymes to break it down Important examples? The biomass at any given trophic level is produced from a much larger biomass ingested from the level below Human Impact on Ecosystems Increasing Carbon Dioxide Concentration in the Atmosphere Burning fossil fuels (wood, coal, oil) releases CO2 Carbon dioxide and water in the atmosphere retain solar heat, causing the greenhouse effect Human Impact on Ecosystems Use of chlorofluorocarbons has destroyed ozone (O3) by converting it to oxygen gas. Ozone protects against UV radiation Increasing skin cancers, cataracts What are your odds of getting skin cancer in your lifetime? Rain Shadow Air rises on the windward side, loses moisture before passing over the mountain Leeward side is in the rainshadow; deserts Figure 49.7 Page 893 Biomes Regions of land characterized by habitat conditions and community structure Distinctive biomes prevail at certain latitudes and elevations Tropical Forests May be dry, deciduous, or rainforests T. Rainforest Abundant rainfall 4 layers to forest (upper & lower canopy, shrub understory, & herbaceous layer) Poor soil due to leaching Highest species diversity Grasslands Savannas Tropical & subtropical with scattered trees 3 seasons: cool & dry; hot & dry; warm wet. Frequent fires Grazing mammals (African grasslands) Chaparral Along coastlines in mid latitudes Mild, rainy winters & hot, dry summers Evergreen shrubs Periodic fires Browsers, rodents reptiles Temperate Grassland (Prairie) Similar to savannah without trees Cold winters Maintained by fire Seasonal drought Rich soils Grazing animals; herbivores Temperate Deciduous Forest Our biome 3 layered forest Dominant species are deciduous trees Midlatitudes Deserts Less than 10 centimeters annual rainfall, high level of evaporation Tend to occur at 30 degrees north and south and in rain shadows One-third of land surface is arid or semiarid Arctic Tundra Occurs at high latitudes Permafrost lies beneath surface Nutrient cycling is very slow Do not post on Internet Coldest biome Arctic tundra in Russia in summer Low species diversity Figure 49.19 Page 903 Taiga (coniferous forest) Found in northern latitudes Harsh winters; short summers Thin, acidic soil Coniferous trees No permafrost Alpine Tundra Occurs at high elevations No underlying permafrost Plants are low cushions or mats as in Arctic tundra Do not post on Internet Figure 49.19 Page 903 Lakes Bodies of standing freshwater Eutrophic: shallow, nutrient-rich, has high primary productivity, Oligotrophic: deep, nutrientpoor, has low primary productivity Lake Zonation LITTORAL LITTORAL LIMNETIC PROFUNDAL Figure 49.21 Page 904 Thermal Layering In temperate-zone lakes, water can form distinct layers during summer THERMOCLINE Figure 49.22 Page 904 Seasonal Overturn In spring and fall, temperatures in the lake become more uniform Oxygen-rich surface waters mix with deeper oxygen-poor layers Nutrients that accumulated at bottom are brought to the surface Ocean Provinces neritic zone oceanic zone intertidal zone continental shelf BENTHIC PROVINCE bathyal shelf PELAGIC PROVINCE 0 200 1,000 abyssal zone 2,000 4,000 hadal zone Figure 49.24 Page 906 deep-sea trenches 11,0000 depth (meters) Phytoplankton Floating or weakly swimming photoautotrophs; form the base for most oceanic food webs Ultraplankton are photosynthetic bacteria Hydrothermal Vents Openings in ocean floor that spew mineral-rich, superheated water Do not post on Internet Primary producers are chemoautotrophic bacteria; use sulfides as energy source Tube worms at hydrothermal vent Figure 49.26 Page 907 Estuary Partially enclosed area where saltwater and freshwater mix Dominated by salt-tolerant plants Examples are Chesapeake Bay, San Francisco Bay, salt marshes of New England Estuarine Food Webs Primary producers are phytoplankton and salttolerant plants Much primary production enters detrital food webs Detritus feeds bacteria, nematodes, snails, crabs, fish Intertidal Zones Littoral zone is submerged only during highest tides of the year Midlittoral zone is regularly submerged and exposed Lower littoral is exposed only during lowest tides of the year Rocky Intertidal Grazing food webs prevail Vertical zonation is readily apparent Diversity is greatest in lower littoral zone Figure 49.29 Page 909 Do not post on Internet Upwelling Upward movement of water along a coast; replaces surface waters that move away from shore Figure 49.31 Page 910