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Ecology of Organisms and Populations Ch. 18 Ecology  Study of interactions between organisms and their environment  Ecology can be divided into four increasingly comprehensive levels:  Organismal ecology  Population ecology  Community ecology  Ecosystem ecology Ecology  Organismal ecology  Is concerned with evolutionary adaptations that enable individual organisms to meet the challenges posed by their abiotic environments. http://www.seabird.org/assets/killer%20whales%20internet%202.jpg Ecology  Population ecology  Is concerned with populations, groups of individuals of the same species living in the same area.  Concentrates mainly on factors that affect population density and growth. http://newsimg.bbc.co.uk/media/images/44609000/jpg/_44609350_puffins512.jpg Ecology  Community ecology  Is concerned with communities, assemblages of populations of different species.  Focuses on how interactions between species affect community structure and organization. http://www.mass.gov/envir/forest/images/multiLayerForest.jpg Ecology  Ecosystem ecology  Is concerned with ecosystems, which include all the abiotic factors in addition to the community of species in a certain area.  Focuses on energy flow and the cycling of chemicals among the various abiotic and biotic factors. http://www.african-books.com/images/Animals/montage.jpg Components of the Environment  The abiotic component  Consists of nonliving chemical and physical factors.  The biotic component  Includes the living factors. Abiotic Factors of the Biosphere  On a global scale, ecologists have recognized striking regional patterns in the distribution of terrestrial and aquatic life.  Global distribution patterns  Reflect regional differences in climate and other abiotic factors. Sunlight  Solar energy powers nearly all ecosystems.  Availability of sunlight affects aquatic and terrestrial environments. http://artfiles.art.com/images/-/Aflo/Sun-Shining-in-BlueSky-Over-Tree-in-Winter-Snow-Biei-Hokkaido-JapanPhotographic-Print-C13062664.jpeg Water  Aquatic organisms may face problems with water balance.  For terrestrial organisms, the main water problem is drying out. Temperature  Environmental temperature  http://www.wildherps.com/images/herps/standard/desert_iguana.jpg http://www.sciam.com/media/inline/A186A7F7-D8EADDDF-0F715313A7DA2A91_1.jpg Is an important abiotic factor because of its effect on metabolism. Wind  Some organisms depend on nutrients blown to them by wind.  Organisms such as plants depend on wind to disperse pollen and seeds.  Can also affect the pattern of a plant’s growth. http://www.asdk12.org/staff/vanarsdale_mark/pages/Ecology_Images/wind_tree.jpg Rocks and Soil  Soil variation contributes to the patchiness we see in terrestrial landscapes.  In streams and rivers, the composition of the soil can affect water chemistry. Periodic Disturbances  Catastrophic disturbances  Can devastate biological communities.  After a disturbance,  An area is recolonized by organisms, or repopulated by survivors.  The structure of the community undergoes a succession of changes during the rebound. Ecosystems  What biotic and abiotic factors do you see in this picture of the rain forest? Ecosystems  What biotic and abiotic factors do you see in this picture of a tundra? Populations • A population is… Members of the same species…  Who live in the same place  At the same time.  Populations • The environment where a population lives: habitat. Populations  Population ecologists study many things about populations in their habitats:  Population size  Population density  Population growth Population Density  Population density  Is the number of individuals of a species per unit of area or volume.   In most cases, it is impractical or impossible to count all individuals in a population. In some cases, population densities are estimated by indirect indicators, such as number of bird nests or rodent burrows. Populations  Populations are densest where there are resources available. Populations Patterns of Dispersion  The dispersion pattern of a population is the way individuals are spaced within the population’s geographic range. Clumped Pattern of Dispersion  Individuals aggregate in patches. Uniform Pattern of Dispersion  Results from interactions among the individuals of a population. Random Pattern of Dispersion  Individuals are spaced in a patternless, unpredictable way. Population Growth Models  Two models, the exponential growth model and the logistic growth model, will help us understand population growth.  The growth rate  Is the change in population size per time interval. The Exponential Growth Model: The Ideal of an Unlimited Environment  The exponential growth model  Describes the rate of expansion of a population under ideal, unregulated conditions. The Exponential Growth Model: The Ideal of an Unlimited Environment • A key feature of the exponential growth model is that the rate at which a population grows depends on the number of individuals already in the population. The Logistic Growth Model: The Reality of a Limited Environment  In nature, a population may grow exponentially for a while, but eventually one or more environmental factors will limit its growth.  Population-limiting factors restrict population growth. Copyright © 2007 Pearson Education Inc., publishing as Pearson Benjamin Cummings The Logistic Growth Model: The Reality of a Limited Environment  The logistic growth model  Describes growth of an idealized population that is slowed by limiting factors.  A comparison of the logistic growth model and the exponential growth model Carrying Capacity  Is the number of individuals in a population that the environment can just maintain with no net increase or decrease. http://www.abc.net.au/reslib/200710/r189329_709751.jpg Regulation of Population Growth  Density-Dependent Factors  Are population-limiting factors whose effects intensify as the population increases in size.  Increase a population’s death rate and decrease the birth rate. Regulation of Population Growth  Density-independent factors  Are population-limiting factors whose intensity is unrelated to population density.  Include events such as seasonal freezing.  In many natural populations, density-independent factors limit population size before densitydependent factors become important. Growth Rate  Four influences:     Birth rate  Death rate  Immigration  Emigration   Birth + Immigration – Death – Emigration Population Cycles  Some populations   Have regular boomand-bust cycles. Boom-and-bust cycles of the snowshoe hare and one of its predators, the lynx Communities and Ecosystems Ch. 19 Key Properties of Communities  Diversity—variety of different kinds of organisms that make it up  Prevalent form of vegetation— determines kinds of organisms that will survive in the area  Stability—ability to resist change and return to its original species composition after being disturbed  Trophic level—feeding relationships among the various species Diversity Which community is more diverse?  The diversity of a community  Is the variety of different kinds of organisms that make up the community.   Species richness, the total number of different species in the community Relative abundance of the different species Interactions Between Populations of Different Species  Interspecific interactions—occur b/w populations of different species  Coevolution—a change in one species acts as a selective force on another species Interspecific Competition  Competition occurs when 2 or more populations overlap in their niches  Limiting resources Food  Space  Mates   Generally, one will out-compete the other Competition in Nature   Two possible Outcomes 1. Weaker competitor becomes extinct 2. One or both species may evolve enough to use a different set of resources (resource partitioning) Competition cannot operate for long periods of time Competitive Exclusion Principle  Two species cannot coexist in a community if their niches are identical Joseph H. Connell Study Interactions Between Populations of Different Species  Predation—consumption of one organism by another  Parasitism—specialized predator (parasite) lives on/in its host, not killed immediately  Endoparasitism—live inside host (tapeworms/viruses)  Ectoparasitism—live on surface of host (mosquitoes/aphids)  Herbivory—herbivores consume plants http://www.dldigital.com/images/z_oldimages/2002-10-d28aphid2-fr18.jpg Predator Adaptations  Most predators have acute senses.  Many predators   Have adaptations such as claws, teeth, fangs, stingers, or poison to help catch and subdue prey. Are fast and agile. http://upload.wikimedia.org/wikipedia/commons/thumb/7/7c/H awk_eating_prey.jpg/300px-Hawk_eating_prey.jpg http://images.encarta.msn.com/xrefmedia/sharemed/targets/images/pho/00123/ 00123ed3.jpg Plant Defenses Against Hebivores www.treklens.com/.../Sweden/phot o198584.htm  Physical defenses  thorns, hooks/spines on leaves   http://en.wikipedia.org/wiki/Image:Toxic odendron_radicans.jpg  Chemical defenses  Make plant distasteful or poisonous Morphine from opium poppy Nicotine from tobacco Poison Ivy http://images.wildmadagascar.org/pictu res/isalo/walking_stick0071.jpg Animal Defenses Against Predators  Behavioral defenses  Alarm cries  Distraction displays  Cryptic coloration/shape (camouflage) Blend in with environment  Asposematic coloration  Red/black; yellow/black Stick Insect www.laspilitas.com/.../Monarch_butte rfly.htm   Mechanical/chemical defenses  Quills, spines, and other similar structures  Toxins—distasteful or poisonous Monarch butterfly on Milkweed Animal Defenses Against Predators  Mimicry—prey resembles species that cannot be eaten  Batesian mimicry: Imitate color patterns or appearance of more dangerous/unpalatable organisms  Müllerian mimicry: 2 unpalatable species that inhabit the same community mimic each other Animal Defenses Against Predators Mimicry can be used to lure  prey  Snapping turtle wriggles tongue like a worm to attract and capture small fish http://www.lancashiremcs.org.uk/g allery/pics/lophius.jpg  Angler Fish attract prey close enough to their mouths to be easily grabbed Symbiotic Relationships  Non-Beneficial  Parasitism—host harmed  Beneficial  Mutualism—both partners benefit Lichens-association b/w fungus and algae  Nitrogen-fixing bacteria and legumes  Community Structure  Predators can moderate competition among its prey species  Keystone species can alter the whole community Community Structure Lake Davis, CA  Introduction of a http://www.dfg.ca.gov/lakedavis/ Northern Pike http://aquanauts_dc.homestead.com/files/northern_pike1.jpg species (exotic species) into a community can have drastic affects on the existing community members Disturbances in a Community  Storms, fire, floods, droughts, overgrazing, or detrimental human activities:  Remove organisms  Alter resource availability  Create opportunities for new species that have not previously occupied the habitat  Humans are the biggest disturbance  Logging, agriculture, overgrazing Ecological Succession  Primary succession  Begins in a virtually lifeless area where soil has not formed  Lichens and mosses colonize first  Soil gradually forms and small plants and shrubs take root  Secondary succession  Occurs where an existing community has been cleared by some disturbance that leaves soil in tact  Earliest plants to recolonize are often those that grow from wind-blown or animal-borne seeds Ecological Succession  Tolerance to abiotic conditions determines early species  Competition among early species shape the succession of an area Mt. St. Helen 1980 Eruption MSH80_st_helens_spirit_lake_before_may_18_1980.jpg http://www.jqjacobs.net/photos/volcano/st_helens.html http://denali.gsfc.nasa.gov/research/volc2/MSHreflection.gif Mt. St. Helen Secondary Succession  http://www.kgw.com/newslocal/stories/L_IMAGE.101688cd0b5.93.88.fa.7c.2791 3b573.jpg Red alder disperses easily and is capable of rapid growth on the nutrientpoor, volcanic deposits.  A red-legged frog –one of the creatures living in one of the dozens of ponds created after the eruption.  70 species of birds, including hummingbirds, western meadowlarks and Savannah sparrows www.kgw.com/news-local/stories/kgw_051505_env... www.kgw.com/news-local/stories/kgw_051505_env... An Overview of Ecosystem Dynamics  An  ecosystem Is a biological community and the abiotic factors with which the community interacts. – Energy flow • Is the passage of energy through the components of the ecosystem. – Chemical cycling • Is the use and reuse of chemical elements within the ecosystem. Energy  Flows through an ecosystem when consumers feed on producers.  Cannot be recycled within an ecosystem, but must flow through continuously. Ecosystem Dynamics  Energy  Depend on the transfer of substances in the feeding relationships, or trophic structure, of an ecosystem.  Trophic  relationships Determine an ecosystem’s routes of energy flow and chemical cycling.  Trophic  flow and chemical cycling levels Divide the species of an ecosystem based on their main sources of nutrition. Trophic Relationships  Ecosystems divided into trophic levels (feeding levels)  Producers—autotrophs (mostly photosynthetic)  Primary consumers—herbivores  Secondary consumers—carnivores that eat herbivores  Tertiary consumers—carnivores that eat other carnivores  Detrivores—consumers that eat dead or decaying matter Food Chain/Food Web Energy Flow in Ecosystems  Each level in a food web contains a different quantity of stored chemical energy  When consumers eat producers or 2 consumers eat 1 consumers, some energy is lost in each transfer from one level to the next Energy pyramid  A diagram that represents the cumulative loss of energy from a food chain. Chemical Cycling in Ecosystems  Ecosystems  Depend on a recycling of chemical elements.  Biogeochemical cycles  Are chemical cycles in an ecosystem that involve both biotic and abiotic components. Biogeochemical Cycles  Three key points :  Each circuit has an abiotic reservoir.  A portion of chemical cycling can rely completely on geological processes.  Some chemicals require processing before they are available as inorganic nutrients. Examples of Biogeochemical Cycles  Carbon  Nitrogen  Phosphorus  Water Carbon Cycle Human Impacts:  Greenhouse Effect   Increase of atmospheric CO2  Combustion of fossil fuels  Burning of wood from deforestation Increase in global temperature Nitrogen Cycle Human Impact:  Cultivation—turns up soil and ↑ decomposition of organic matter; Releases more nitrogen  Harvesting ↓ nitrogen from ecosystem  Adding industrially synthesized fertilizers to soil has resulted in doubling globe’s supply  Excess nitrogen leeches into soil and into rivers, streams, and lakes and ground water— – ↑ levels are toxic to aquatic organisms and humans – Algal blooms in lakes ↑ eutrophication Phosphorus Cycle Human Impact:  Sewage treatment facilities and fertilizers  ↑ amounts of phosphates to aquatic systems, causing eutrophication of lakes. Water Cycle Human Impact:  Destruction of tropical rain forest   Will change the amount of water vapor in the air. May alter local and global weather patterns.  To irrigate crops, humans pump large amounts of ground water to the surface.
 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                            