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Transcript
Dynamics of Ecosystems
Chapter 57
1
Flow of Energy in Ecosystems
• First Law of Thermodynamics: energy is
neither created nor destroyed; it changes
forms
• Second Law of Thermodynamics: whenever
organisms use chemical-bond or light energy
some is converted to heat (entropy)
• Sun our major source of energy (E)
2
Flow of Energy in Ecosystems
• Trophic levels: level an organism “feeds” at
• Producers (autotrophs): “self-feeders” make
organic compounds (photosynthesis)
• Consumers (heterotrophs): must take in food
3
Flow of Energy in Ecosystems
• Consumers are classified by their diet
• Herbivores: first consumer level, eat plants
• Primary carnivores: eat herbivores
• Secondary carnivores: eat primary
carnivores or herbivores
• Detritivores: eat decaying matter
–Decomposers: microbes that break up
dead matter – “CHONPS”
4
Trophic levels within an ecosystem
5
Flow of Energy in Ecosystems
• Productivity: the rate at which the organisms in the
trophic level collectively synthesize new organic
matter
• Primary productivity: producers
• Respiration: rate producers use org. compounds
• Net primary productivity (NPP) = PP – respiration
• Secondary productivity: productivity of a
heterotroph trophic level
6
Flow of Energy in Ecosystems
• biomass: the amount of organic matter
present at a particular time
• Only small fraction of incoming solar
energy is captured by producers ~ 1%/year
– Used to make chemical-bond energy
– As energy passes up the food chain, most is
lost as heat and waste
– Less biomass/fewer individuals at each
trophic level
7
Flow of Energy in Ecosystems
• 50% of chemical-bond energy is not assimilated
and is egested in feces
• 33% of ingested energy is used for cellular
respiration
• 17% ingested energy is converted into insect
biomass
8
9
Flow of Energy in Ecosystems
Flow of energy through the trophic levels of Cayuga Lake
10
Human Impacts:
Pollution
• Biomagnification:
becomes more
concentrated at
higher trophic levels
• predatory bird
species’ eggshells so
thin that the shells
broke during
incubation
11
12
Ecosystem productivity per year
13
Flow of Energy in Ecosystems
• Trophic level interactions
– Trophic cascade: process by which effects exerted
at an upper level flow down to influence two or
more lower levels
– Top-down effects: when effects flow down
– Bottom-up effects: when effect flows up through
a trophic chain
14
Flow of Energy in Ecosystems
Trophic cascade in a large-scale ecosystem
15
Example: Top-down
• Human removal of carnivores produces topdown effects
– Over fishing of cod - 10% their previous numbers
– Jaguars and mountain lions absent on Barro
Colorado Island
– Smaller predators become abundant
16
Example: bottom-up
• When primary productivity is low, producer
populations cannot support herbivore
populations
• As primary productivity increases, herbivore
populations increase
• Increased herbivore populations lead to
carnivore populations increasing
17
Flow of Energy in Ecosystems
Bottom up effects
18
Biodiversity and Stability
• Species richness is influenced by ecosystem
characteristics
– Primary productivity
– Habitat heterogeneity
• Accommodate more species
– Climatic factors
19
Biodiversity and Stability
Factors that affect species richness
20
Biodiversity and Stability
• Tropical regions have the highest diversity
–Species diversity cline: biogeographic
gradient in number of species correlated
with latitude
– Evolutionary age of tropical regions
– Increased productivity
– Stability/constancy of conditions
– Predation
– Spatial heterogeneity
21
Biodiversity and Stability
Latitudinal cline in species richness
22