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Variations are inherited from one generation to the next leading to natural selection Differences that help organisms survive and reproduce become more common and differences that are not beneficial become less common It changes the genes!!! Darwin did not know about how things are inherited yet (Mendel). We can use our knowledge from Mendel’s study of heredity and combine it with Darwin’s study of evolution to explain how inheritable variation appears and how natural selection effects that variation. • • Biologists studying evolution focus on a population = collection of individuals of the same species in a given area A population shares a common group of genes (genetic information) = gene pool – • Contain two or more alleles (forms of a gene) for each inheritable trait Relative frequency (%) = the number of times an allele occurs in a gene pool compared to the number of times other alleles occur Section 16-1 Sample Population 48% heterozygous black 36% homozygous brown 16% homozygous black Frequency of Alleles allele for brown fur allele for black fur • Mutations = any change in a sequence of DNA Remember: mutations result as a mistake during replication or toxin (chemicals/radiation) – Some mutations effect phenotypes (physical characteristics), which can effect an organism’s fitness (ability to survive) – • Gene shuffling = different gene combinations inherited during gamete production creating different genotypes (genetic makeup), different phenotypes and more variation Crossing over increases number of different genotypes – Does not change the relative frequency of alleles in a population – – If it is a trait controlled by a single-gene with 2 alleles, there will be only 2 phenotypes. • – Frequency of Phenotype (%) The number of phenotypes produced for a given trait depends on how many genes control the trait. Phenotypes of a single-gene is represented by a bar graph If it is a trait is polygenic with 2 or more alleles, there will be many genotypes and even more phenotypes. • Phenotype Phenotypes of a polygenic trait is represented by a normal distribution (bell curve) Frequency of Phenotype • Phenotype (height) Natural selection acts on phenotypes, survival and reproduction determine which alleles are inherited, changing relative frequencies of alleles in a population over time. Thus evolution is any change in the relative frequencies of alleles in a population’s gene pool and acts on populations, not individuals. Natural selection on single-gene traits can lead to changes in allele frequencies and thus to evolution. One of the two phenotypes may make an organisms better fit, thus under pressure from natural selection and its relative frequency will increase • Natural selection can affect the distributions of phenotypes in any of three ways: – – – Directional selection: When the entire bell moves left/right because there’s a higher fitness and increase in the number of individuals with the trait at one end of the curve Stabilizing selection: When the bell becomes more narrow, because there’s a higher fitness and increase in the number of individuals with the trait in the center of the curve Disruptive selection: The bell can split into two, because there’s a higher fitness and increase in the number of individuals at both ends of the curve Section 16-2 Key Directional Selection Food becomes scarce. Low mortality, high fitness High mortality, low fitness Section 16-2 Stabilizing Selection Key Low mortality, high fitness High mortality, low fitness Birth Weight Selection against both extremes keep curve narrow and in same place. Section 16-2 Disruptive Selection Low mortality, high fitness High mortality, low fitness Population splits into two subgroups specializing in different seeds. Beak Size Number of Birds in Population Key Number of Birds in Population Largest and smallest seeds become more common. Beak Size • • • • Genetic drift = random change in allele frequencies In small populations, individuals that carry a particular allele may leave more descendants than other individual, just by chance. Over time, a series of chance occurrences can cause an allele to become common in a population. Genetic drift can happen when a small group of individuals colonize a new habitat carrying different relative frequencies that the larger population. Founder effect = allele frequencies change as a result of the migration of a small subgroup of a population Section 16-2 Sample of Original Population Descendants Founding Population A Founding Population B Section 16-2 Sample of Original Population Descendants Founding Population A Founding Population B Section 16-2 Sample of Original Population Descendants Founding Population A Founding Population B The conditions that must be met to avoid evolution: Random mating Large population No movement into or out of the population No mutations No natural selection • • • Speciation = formation of new species Species = group of organisms that breed with one another and produce fertile offspring (share a common gene pool) As new species evolve, populations become reproductively isolated from each other: When 2 populations can’t breed and produce fertile offspring, resulting in separate gene pools Behavioral isolation: Capable of breeding, but have different courtship rituals or behaviors – Geographic isolation: Separate by geographic barriers – Temporal isolation: Reproduce at different times – • Speciation in the Galapagos finches occurred by: – – – – – – Founding a new population: A small population of finches migrates to a different island Geographic isolation: Finches don’t usually fly over open water, so stayed on own island (separate gene pool) Changes in the new population’s gene pool: Adapted to new environment (directional selection) to be more fit Reproductive isolation: Differences in phenotypes and mating rituals may turn different finches off to one another Ecological competition: Similar finches compete, so individuals that are most different from each other have the highest fitness, because less competition. Continued Evolution: Process repeats and over many generations, it produced the 13 different finch species