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Phylogeny and Systematics The Tree of Life Phylogeny is the evolutionary history of a species (or group of species) To determine phylogeny, evolutionary biologists use: The fossil record Systematics An analytical approach to understanding relationships between organisms Traditionally have used morphological and biochemical resemblances (homologous structures, etc.) ***Molecular systematics – Compares DNA and RNA to infer evolutionary relationships The Fossil Record The fossil record is based on the sequences in which fossils have accumulated in strata of rocks Relative vs. absolute dating The fossil record is an important but incomplete chronicle of evolutionary change Only organisms with hard parts (ie. skeleton) could be preserved as fossils Many fossils have not been discovered Morphological and Molecular Homologies In general, organisms that share very similar morphologies (forms) or similar DNA sequences are more likely to be closely related Homologous structures Examples: Convergent Evolution HOWEVER, convergent evolution occurs when similar environmental pressures and natural selection produce similar (analogous) adaptations in organisms from different evolutionary lineages In other words, the SAME PRESSURES cause the SAME adaptations to evolve INDEPENDENTLY Example: Bats and birds have adaptations that allow them to fly This might imply that bats and birds are closely related, perhaps more closely related than bats and birds Bat’s forelimb is homologous to other mammals, but analogous to a bird’s wing Homologous vs. Analogous Structures Homologous Structures – Similar anatomical structures that came from a common ancestor as a result of divergent evolution Analogous Structures – Anatomical structures that are similar and perform the same/similar function but evolved SEPARATELY A result of convergent evolution Taxonomy Taxonomy is the ordered division of organisms into categories based on similar characteristics Carolus Linnaeus began taxonomy in the 18th century However, his system of taxonomy is under some attack now because his classification was based on resemblances, NOT evolutionary relationships Linnean Classification Binomial Nomenclature: 2-part format of the scientific name Genus and species Hierarchical Classification Linnaeus grouped species into a hierarchy of increasingly broad categories Genus – Family – Order – Class – Phylum – Kingdom – Domain Apt. - Apt. Bldg. – Street – City – State - Country - Continent Phylogenetic Trees Phylogenetic trees are branching diagrams that show hypotheses about evolutionary relationships between organisms REMINDER: Phylogenetic trees represent HYPOTHESES about how organisms are related A good hypothesis must be revised when new data is revealed Cladograms Cladograms show patterns of shared characteristics and can help us infer evolutionary relationships (as long as they’re due ONLY to homology, NOT analogy)