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Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill UNIT B: Ecosystems and Population Change General Outcome 1: Explain that the biosphere is composed of ecosystems, each with distinctive biotic and abiotic characteristics. Cell Division: Mitosis Asexual cell division Body (somatic) cells divide to form two identical daughter cells. o The DNA is first replicated (copied or doubled) o Then the cell divides, with a copy of the DNA in each new cell. Remember: DNA is organized in chromosomes. Individuals have one set of chromosomes from their mother, and one set from their father Example: humans have 46 chromosomes – 23 from their mother and 23 from their father or 23 PAIRS of chromosomes Cell Reproduction: Meiosis Sexual cell division Gamete cells (sex cells – egg or sperm cells) pass through two divisions in order to produce sperm or egg cells with half the DNA o First division – one of each pair of chromosomes moves into each new cell Only 23 chromosomes in each new cell (X) o Second division – the new cells (with 23 chromosomes) divide again. Each chromosome (X) is pulled in half resulting in 4 gametes Each gamete has 23 single strands of DNA 1 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Section 3.1: Individuals, Populations, and Communities in Ecosystems Western science has divided the physical world into living (biotic) and non-living (abiotic) components in order to describe and explain organisms and their interactions. Other cultures have different ways of organizing the physical world. FNMI (North American Aboriginal peoples) consider all the physical world to be living. o Have organized the material world into 4 directions (E, N, W, S) with specific characteristics. Ancient Greeks debated over what was living or non-living. o Also divided the physical world into 4 components: Earth, Air, Fire, and Water Environment – everything that affects an organism throughout its life, as well as everything that the organism affects. Biotic – (Living) bacteria, micro-organisms, insects, parasites, predators, prey, and the interactions between living things Abiotic – (Non-living) sunlight, water, minerals (nutrients), heat, weather conditions, pH, soil, availability of shelter, wind, Ecologists – scientists who study the interactions of organisms with one another and their environment. Levels of the biotic environment: 1. 2. 3. 4. 5. Organisms Populations Communities Ecosystems Biosphere Note: smaller than organisms are individual cells, larger than biosphere is the solar system… Organisms – individual organisms. Scientists study the individuals abiotic environment to learn how it affects the individuals behavior or physical features or distribution May study the physiology (physical features) of the organism – specifically adaptations that allow it to live/survive in a particular environment. Abiotic conditions affect the distribution of organisms. Populations – a group of individuals of the same species living in a specific area at the same time. Species – organisms that are able to breed with one another and produce fertile offspring. 2 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Population ecologists study: o Changes in the size of populations over time o The distribution (abundance) of populations o Rate of population change in size o Gender and age distributions in the population o Factors (biotic or abiotic) affecting the population Communities – all of the individuals in all of the interacting populations in a given area. Community ecologists study: o Interactions between different populations in the community Competition Intraspecific – competition within a population o Competition for mates o Competition for food o Density dependent Interspecific – competition between different populations Predator/prey o Parasitism o Diversity of communities within different environments o Abiotic factor affecting different communities 3 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Most communities are dynamic – they change continually over time o As abiotic conditions change Seasonally In response to human interaction Global weather fluctuations o As populations fluctuate Intraspecific competition Interspecific competition Predator-prey relationships Over hunting or over-fishing Competition results in stress which can lead to: Disease Lower body size Lower birth rate Lower birth weight Reduced fertility Changes in reproductive readiness times between males and females Higher availability of food can cause increases in populations Loss of a predator Greater input of nutrients into the ecosystem o As the environment changes forest fires or prairie fires land slides rock slides volcanic activity glacier recession 4 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Ecosystems – a community of populations, together with the abiotic factors that surround and affect it. The size of the ecosystem depends upon what you are studying. (Remember, ecosystems are human constructs, just a description of what we want to investigate.) Biosphere – all the ecosystems in the world; the largest possible ecosystem. Populations are not randomly scattered throughout the biosphere. Species distribution is dependent on the availability of biotic and abiotic factors (food, shelter, sunlight, moisture, etc.) Section 3.2: Classifying and Naming Organisms When we investigate natural systems, we need to clearly communicate our findings to others (scientists, media, lay-persons). Taxonomy – the practice of classifying living things. Aristotle (Greek philosopher) – attempted to classify all known life (1000 different organisms) o Two main groups – plants and animals – each group called Kingdoms. o Categorized animals based on their size and the way they moved on land, in water, or in air. o Plants were categorized into three groups based on differences in their stems. Other philosophers and naturalists used different criteria for categorizing living things o Structures o Reproductive structures o External tissues Ernst Haeckel (1866) – after the discovery of the microscope o Classified micro-organisms based on feeding Photosynthetic or non-photosynthetic Some micro-organisms have both processes grouped into a third kingdom called Protista 5 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Classification is still changing – due to the findings of biochemistry we now have a level of classification above kingdom domain Domain Kingdom Phylum Class Order Family Genus Species Bacteria Eubacteria Prokaryotic Asexual Heterotropic Autotrophic Flagella cyanobacteia Archaea Archebacteria Prokaryotic Asexual Protista Eukaryotic Single or multiple celled Chemosynthesis or heterotrophic Distinctive chemistry Sexual repro Eukarya Fungi Plantae Eukaryotic consumer by absorption no photosynthesis Eukaryotic Photosynthetic Cell wall Animalia Eukaryotic Cell membrane Mobile Consumers Amoeba algae The classification system moves from large, general categories into successively narrow, more specific categories, down to species. Example: Humans Domaine – Eukarya Kindom – Animalia Phylum – Chordata Class – Mammalia Order – Primates Family – Hominidae Genus – Homo Species – Sapiens 6 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Carolus Linnaeus (1707 – 1778) – Swedish biologist Subdivided each kingdom into smaller and smaller groups of more and more similar organisms o Used physical characteristics o At each level there are more similarities among members of the group Used the naming system to devise a simple, scientific name for organisms o Binomial Nomenclature (2 name naming system) Used the genus and species names. Genus – capitalized. Species – lowercase Written in italics or underlined if handwriting. Can shorten the genus name to the capital letter. Example: Homo sapiens or H. sapiens Puma concolor = North American cougar Felix domesticus = common house cat o To make the naming system universal, we use Latin, a language spoken in no country, but which (at the time) all educated people understood, and which forms the basis for many of the Western languages (English, German, French, Spanish, Portugese, Italian) Dichotomous Keys Identification key using observable characteristics (or paired comparisons) to identify known organisms (already classified organisms). These are arranged in steps – each step is an ‘either/or’ question. Example: red or not red. 1. Select one characteristic, then sort the items/organisms into two groups based on whether they have this characteristic. 2. Select a second characteristic for each subgroup, and sort. 3. Continue to subdivide the groups until you have sorted each item/organism. 7 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Section 3.3: Studying Organisms in Ecosystems Climate – the average weather conditions in a particular region over a period of time, usually 30 years or more. - - Determined by rainfall, which results from the unequal heating / cooling or Earth, local geography (topography, altitude, latitude), snow/ice cover, and the proximity of large bodies of water. The pattern of precipitation influences the type of soil that forms in different regions. Producer Distribution - - Plant types/abundance is determined by: o Moisture levels o Heat - temperature o Types of soil o Topography o Altitude o Latitude o Abundance/intensi ty of sunlight The types of plants determines the variety and population sizes of non-photosynthetic organisms (consumers – fungi, decomposers, animals) Biomes – very large ecosystems or groups of ecosystems with a particular mix of plants, animals, and other organisms that are adapted to living under the specific environmental (abiotic) conditions within that region. - May be terrestrial or aquatic biomes 8 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill - Terrestrial biomes – the abundance of life increases as the temperature and precipitation increase o Temperature changes with latitude and altitude, resulting in biome patterns - Aquatic biomes – the abundance of life increases with sunlight and nutrient availability o Ocean surface – sunlight availability o Continental shelf – sunlight and nutrient availability There are gradual transitions between biomes boundaries resulting in a gradual change in the composition of plants and animals - 9 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Habitats – (home) a place or area with a particular set of characteristics, both biotic and abiotic. Organisms within that habitat are specifically adapted to those characteristics. - Habitat may spread over a large area, or be found in different locations (ex. Alpine meadows) Range – the geographical area where a population (species) is found. - Range is determined by habitat requirements. The species will only be found where the habitat is present. The organism may not live throughout the entire range, but only in its particular habitat within the range. 10 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Niches – (job) the role of a species in a community and the total range of biotic and abiotic requirements that the species need in order to survive. - - Each species has its own specific niche. No two species can share the same niche. o If two species occupy the same niche, they will compete for resources, and one or both will be eliminated. Niches may be similar (overlapping) but not identical in order for different species to coexist. Aquatic Habitats and Niches - - Aquatic ecosystems have vertical stratification (layering) of water caused by different physical and chemical conditions. o Sunlight levels o Nutrient levels o Temperature levels o Oxygen levels Each species in a lake occupies a niche that it requires for survival. 11 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Biodiversity – the greater the variety of habitats and niches, the greater the variety of species. Limiting Factors – abiotic or biotic factors that limit a populations growth. - Abiotic limiting factors – soil types, pH, nutrient requirements, moisture, weather, temperature 12 - Biodiversity varies widely depending on the specific abiotic and biotic components of the ecosystem. - Monocultures – (replanted forests, agriculture fields, domestic lawns) show a reduction in habitat diversity, resulting in a reduction of biodiversity (due to loss of habitat) Biology 20 - Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Biotic limiting factors – competition, limited nutrients, predators, parasites, introduced (exotic) species 13 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Sampling Populations – provides information on types of species, their density, and changes in populations and communities - A census is too time consuming, and often impractical Taking samples – small portions or subsets of the entire population and then calculating the average is easier o Transect – lay a line across the area you wish to sample. May count only organisms touching the line, or may count any organisms within a set distance from the line. Calculate the total area Divide the number of organisms by the sample area o Quadrats – a premeasured area (usually 1 m2) Several locations within the study area are chosen (random) At each location a quadrat is marked out The number of individuals within the quadrat are counted The density (number of organisms divided by the area) is determined by taking the average of the quadrats. 14 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill General Outcome 2: Explain the mechanisms involved in the change of populations over time. Section 4.1: Adaptation, Variation, and Natural Selection No two organisms, even within the same species, are identical. Differences may be physical (observable) – colour, size, or physiological (metabolic functioning, not directly observable) – disease resistance, ability to tolerate stress. These differences are examples of variation within a population, and help (or hinder) organisms to survive within their specific habitat and environmental conditions. Variation within a population is the result of sexual reproduction. Each sperm or egg produced contains a different sampling of the parents DNA, so each joining of a sperm and egg produces slightly different combinations. Organisms that survive long enough to reproduce have the opportunity to pass along to their offspring the genetic information that helped them survive. The more an individual reproduces the more chances that successful genetic material will be passed along. Adaptation – the specific biochemical/physical/behavioral characteristics of an organism that helps it survive within its environment. - Adaptations are the result of a gradual change in the characteristic of members in a population over time. Structural adaptations – examples: Human – opposable thumb, pelvic structure that enables bipedalism, binocular vision, development of the frontal cortex in the brain, Biochemical adaptations – examples: Human – inability to synthesize vitamin C (ingestion only), insulin and energy storage within the body, female reproductive hormones Behavioral adaptations – examples: Human – the need for human contact/relationships Variation – a visible or invisible difference in the genetic/physical/behavioral make up of an individual. - May either be neutral (have no effect), harmful (reduce an individual’s chance to pass the trait along to its offspring), or beneficial (help an individual’s chance to pass along traits to offspring). 15 Biology 20 - Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Not all variations become adaptations Biological Species – a group of reproductively compatible populations. - - Members of these populations can interbreed and produce viable (able to reproduce) offspring. Offspring have a combination of genetic material from both parents through sexual reproduction. o The number of possible variations in genes that offspring can receive is the basis behind variation among individuals. Genetic variation in a population is due to the total genetic variation of all the individuals within that population. Mutations – changes in the genetic material (DNA) of an organism. - - - Mutations happen continuously o Spontaneously during copying of DNA in cell division o Mutagens – environmental agents that can cause mutations in DNA. UV radiation, radiation, smoking, pesticides, etc. When DNA mutates, the cell carrying the DNA may show new characteristics. o Example: a normal rbc versus sickle cell rbc o Cells may not work properly o Cells may die o Cells may begin to divide rapidly (cancer) If the mutated DNA occurs in a body cell (somatic cell), the changes will not be passed along to offspring. If the mutated DNA occurs in a sex cell (gamete – egg or sperm) the changes may be passed on to succeeding generations. Mutations are a significant source of genetic variation in populations. 16 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Selective Advantage – when a mutation enables an organism to better survive in its environment – and thus more likely to pass on that trait to offspring. - - This is more common when an organism’s environment is changing – mutations that were silent (no advantage or maybe a slight disadvantage) are now favorable, allowing the individual organism to survive better. o Examples: housefly’s that are able to resist the pesticide DDT Mutation present before the invention of DDT – caused fly’s to have a slower growth rate a disadvantage. This mutation provided resistance to DDT, allowing these fly’s to survive the pesticide and produce offspring. In populations that reproduce quickly (like bacteria) a rapidly changing environment can lead to rapid adaptation to the change. Natural Selection – the process that results when the characteristics of a population of organisms change because individuals with certain inherited traits survive specific local environmental conditions and, through reproduction, pass on their traits to their offspring. - Requirements for Natural Selection to Occur: o There must be (genetic) variation within the population. o The environment selects for helpful traits – environmental pressure may select FOR some traits or select AGAINST others Selective Pressure o Individuals do not change during their lifetime – the population changed over generations through the accumulation of specific genetic traits/variations o Situational – adaptations to a changing environment are not anticipated or predicted by the individuals or populations…rather the variation is already present, and IF it is helpful WHEN the environment changes, then the individual may survive better. Adaptation to one environment may be harmful in another environment. Example: human energy storage for lean times leads to obesity in times of excess food. 17 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Section 4.2: Developing a Theory to Explain Change - Scientific knowledge develops as people continue to ask questions and search for new answers. Scientific Hypothesis – a statement that provides one possible explanation for an observation or question. - Hypothesis are tested (peer reviewed journals) Explanations that remain useful after new information is found AND which provide predictions that later turn out to be accurate are successful and stay in use for long periods of time. Scientific Theory – a successful hypothesis that makes accurate predictions about a broad range of observations. Example: the Theory of Evolution by Natural Selection – well supported, widely accepted (evidence to back it up…NOT PROOF!), and able to make predictions…therefore science uses this theory until a better theory is developed. 18 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Plato (427-347 BCE) and his pupil Aristotle (384 – 322 BCE) - Believed all life existed in a perfected and unchanging form. This belief was unchanged until after the Renaissance – the movement towards nature as the ideal and observation the means to study it. Buffon (1707 – 1788) – published Histoire Naturelle (44 volumes!) - Noted the similarities between humans and apes Speculated they might have a common ancestor Suggested the Earth was older than commonly thought (older than 6000 years) Cuvier (1769 – 1832) – developed the field of paleontology (study of ancient life through the examination of fossils) - Each stratum (layer) of rock contains specific groups of fossil species Deeper stratum contain older fossils Found that over time (through the layers of rock) some species disappeared and new species appeared – evidence that species could become extinct Proposed that Earth had experienced many destructive events 19 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Lyell (1797 – 1875) – Scottish geologist - Rejected the idea of catastrophic events in Earths past. Proposed that geological processes operated at the same rates in the past that they do today. Slow processes over a long period of time could result in substantial change Inspired Charles Darwin…if the earth could change slowly over time, could change also occur in populations? Lamarck (1744 – 1829) – Philosophie Zoologique - Outlined his theory of how changes occur in a species over time. Compared current species with fossil forms Observed a ‘line of descent’ or progression o A series of fossils (from older to more recent) led to modern species Thought species increased in complexity over time - until reaching a level of perfection. Believed traits (characteristics/variation) was acquired during an organism’s lifetime which could be passed on to offspring. Theory of Inheritance of Acquired Characteristics o While not accurate, he did provide the first theory for inheritance of traits, and that traits could be passed on to offspring. 20 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Darwin (and Wallace ) - - 1809 – 1882 HMS Beagle – naturalist on a multi-year tour of South America Explored the natural history / environment of many countries/geographical locations Didn’t always understand the significance of what he was seeing, but he documented everything so was able to review the information later. Wallace did a similar tour and found similar findings. Communicated with each other, and presented their findings in 1858. o Both accepted that populations change over time o Both not sure HOW populations changed. Malthus (1766-1834) - An economist wrote “Essay on the Principles of Population” Proposed that populations produced far more offspring than their environment could support (food supply) o Populations were eventually reduced by starvation and/or disease o This idea provided Darwin/Wallace with the key Natural Selection - - Individuals have physical, behavioral, biochemical traits which help them survive in their environment. o Those with the traits that helped them survive long enough to produce the most (and healthiest) offspring were more likely to pass on those traits. Competition for limited resources among the individuals WITHIN the same species selects for successful traits Nature provides the pressure for the selection process (resources, food, etc) Present forms of life have arisen by descent and modification from an ancestral species Modification is done by natural selection over long periods of time o All life had descended from some unknown organisms 21 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill o As descendents spread out over different habitats the new environments ‘selected’ for different traits 22 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Evidence of Evolution Fossil Record – sedimentary rock with fossils o Fossils found in young layers of rock are much more similar to species alive today than fossils found in deeper/older layers of rock o Fossils appear in chronological order in the rock layers o Not all organisms appear in the fossil record at the same time Amphibians Mammals Birds Reptiles All appear at different ‘times’ or layers o Transitional fossils – fossils that show intermediary links between groups of fossil organisms and recent organisms Biogeography – the study of the past and present geographical distribution of organisms o Geographically close environments are more likely to be populated by related species than are locations that are geographically separate but environmentally similar Cacti are found in north, central, south America, but not in Australia or Africa o Animals found on islands often closely resemble animals found on the closest continent Suggests that island animals are evolved/descended from mainland migrants o Fossils of the same species can be found on the coastline of neighbouring continents (Africa and south America) Continental drift Pangea 23 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Anatomy – vertebrate skeletal structures (forelimbs, pelvic structure, etc) can be compared between forms o Forelimbs – can be used for running, flying, swimming, swinging All vertebrate forelimbs contain the same set of bones, organized in similar ways Homologous Structures – similar structure but different uses Inherited from a common ancestor Analogous Structures – different structures but similar functions ex. Bat wing, bird wing, insect wing…. Embryology – embryos of different (but related) organisms show similar stages of embryonic development. o Vertebrates – all vertebrate embryos have similar patterns of embryogical development 24 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Molecular Biology and Genetics– studying the molecular/atomic structures of organic compounds (DNA, protein, etc) - All cells consist of membranes filled with water, DNA, proteins, lipids, and carbohydrates Proteins called enzymes control biochemical reactions in all organisms In all organisms, proteins are assembled from amino acids All cells that can replicate contain DNA o Comparing the organization of DNA can indicate how closely related two species are (more differences means more mutations over time, which means longer time as a separate species) o All DNA is made up of the same 4 chemicals Section 4.3: How Species Form Species – a population that can interbreed and produce a group of viable offspring, which can also reproduce in nature. - Can be described as being reproductively isolated from other species Timing of reproductive cycles Arrangement of reproductive organs Separation by geographic barriers 25 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill Speciation - the formation of a new species, resulting from accumulated changes in the population over time. 1. Transformation – when the new species gradually replaces the old species 2. Divergence - One or more species may arise from a parent species that continues to exist. Geographical Barriers – Mountains Rivers Landslides Island formation o These barriers must be present for a long enough time for the populations to become reproductively incompatible with the original population. o Some species may live in the same general area but occupy different niches so don’t encounter each other Biological Barriers – o Behavioral barriers courtship songs Courtship colouring/plumage/displays o Chemical barriers Pheromones for mate attraction Utilization of different resources than the parent population occupation of a new niche (enzymes to take advantage of the new resource?) Adaptive Radiation – the diversification of a common ancestral species into a variety of species, all of which are differently adapted. 26 Biology 20 Unit 2: Ecosystems and Population Change Tannant/McGraw-Hill The Pace of Evolution 1. Gradualism – gradual changes occur steadily, in a linear fashion 2. Punctuated Equilibrium – rapid change after a period of mass extinction, but little change otherwise a. Most physical/morphological change occurs with the first divergence from the parent species Requirements for Speciation by Natural Selection 1. Life forms have developed from ancestral species. 2. All living things are related to one another by varying degrees through common descent. 3. All living things on Earth have a common origin (share a common ancestor). 4. The mechanism by which one species evolves into another species involves random heritable genetic mutations. a. Some mutations result in a survival advantage for an individual; if so that individual is more likely to survive and pass this mutation on to its offspring. Eventually this successful mutation increases in the population and causes the populations as a whole to start to change. 27