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The Sensory Organs Liu jiao Binzhou Medical University Department Of Anatomy The Sensory Organs The sense organs are actually extensions of the nervous system that respond to changes in the internal and external environment and transmit action potentials (nerve impulses) to the brain. It is through the sense organs that we achieve awareness of the environment, and for this reason they have been described as “windows for the brain.” The Sensory Organs A stimulus must first be received before the sensation can be interpreted in the brain and the necessary body adjustments made. A sensation is an awareness of a bodily state or condition that occurs whenever a sensory impulse is transmitted to the brain. The interpretation of a sensation is referred to as perception. Perceptions are the creations of our brain; in other words,we see, hear, taste, and smell with our brain. The Sensory Organs In order to perceive a sensation, the following conditions are necessary. A stimulus sufficient to initiate a response in the nervous system must be present. • A receptor must convert the stimulus to a nerve impulse. A receptor is a specialized peripheral dendritic ending of a sensory nerve fiber or the specialized receptor cells associated with it. • The conduction of the nerve impulse must occur from the receptor to the brain along a nervous pathway. • The interpretation of the impulse in the form of a perception must occur within a specific portion of the brain. The Sensory Organs our perceptions allow us to interact effectively with our environment and are of obvious survival value. The Sensory Organs Sensory organs include the receptors and accessory organs. The receptors may be divided into three kinds: The exteroceptors: receive stimuli such as touch, temperature, pain, light and sound from the external environment The interoceptors: pick up information about internal environment The proprioceptors: Proprioception means "sense of self". In the limbs, the proprioceptors are sensors that provide information about joint angle, muscle length, and muscle tension, which is integrated to give information about the position of the limb in space. The Visual Organ 视器 Composition: eyeball and accessory organs of eye Shape of eyeball Has anterior and posterior poles Equator: an imaginary line encircling the eyeball, midway between anterior and posterior poles Axis of eyeball: a line joining the two poles Optic axis: a line joining the center of the pupil to the fovea centralis Walls of eyeball Cornea Fibrous tunic of eyeball Sclera Iris Vascular tunic of eyeball Cilliary body Choroid Pars iridica retinae Retina Pars caeca retinae Pars ciliaris retinae Pars optica retinae Fibrous tunic of eyeball (outer) Cornea : anterior 1/6, a nonvascular structure, transparent portion, richly supplied by nerves; because it is curved, the cornea helps focus light Sclera (white of eye, opaque portion): posterior 5/6, consisting of fibrous connective tissue that forms a tough protective covering for eyeball, Contains sinus venosus sclerae which lies beneath the junction of cornea and sclera Vascular tunic of eyeball (middle) Iris Thin contractile membrane anterior to ciliary body, with a central opening, the pupil Contains sphincter pupillae (circular fibers) and dilator pupillae (radial fibers) Cornea and iris meet to form the iridocorneal angle Cilliary body Body a ring-shaped thickening anterior to equator, containing smooth muscle fibers called ciliary muscle Ciliary processes : a series of some 60~80 projections producing aqueous humor Ciliary ring Sinus venosus sclerae Ciliary Muscle Iridocorneal angle Dilator Pupillae Sphincter Pupillae Lens ciliary zonule Ciliary Processes Ciliary ring Choroid Thin, highly vascular in posterior 2/3 of eye Contains brown pigmented cells and dense capillary plexus Retina It is the neural sensory stratum lining the inner surface of the vascular tunic Pars caeca retinae Pars iridica retinae Pars ciliaris retinae Pars optica retinae Lines the choroidsComposed of two layers An outer pigment cell layer Inner neural layer (four layers) The fourth layer consists photoreceptor cells Cone cells are color receptors that function best during the day Rod cells are dark-light receptors that function best at night and in dim light The third layer consists of bipolar neurons The second layer is formed ganglion cells, whose axons form optic nerve The first layer consists of nerve axons that collect at the optic disk and pass through the sclera to form the optic nerve Ganglion cells Bipolar neurons Rod cells Cone cells Pigment cell layer Optic disc (blind spot), located medial to posterior pole of eye, and consists of optic nerve fibers and central artery of retina Macula lutea Lies lateral about 3.5 mm to optic disc, a shallow depression, it is completely free of blood vessels and is yellowish in color Fovea centralis, aera of greatest visual acuity (concentration of cones), at its center The pigmentted layer absorbs light that passes completely through the anterior layer, preventing backscatter (blurring of vision) Contents of eyeball Aqueous humor Lens Vitreous body Aqueous humor Chamber of eye - lies between cornea and lens, and divided by iris into anterior and posterior chambers Aqueous humor A clear watery fluid that fills chamber of eye Continuously secreted by ciliary body into posterior chamber Passes through pupil into anterior chamber Then it filters though iridocorneal angle into sinus venosus sclerae, this sinus drains via anterior ciliary veins into ophthalmic veins Production and circulation of aqueous humor Ciliary body Posterior chamber Iridocorneal angle Pupil Sinus venosus sclera Ophthalmic vein Functions • Helps focus light • Helps maintain constant pressure in eyeball • Helps nourish the lens and cornea Anterior chamber Anterior ciliary vein Lens Transparent biconvex structure, covered by an elastic transparent capsule Located between iris and vitreous body, and suspended behind pupil by ciliary zonule Shape changed by the ciliary muscle: for near vision, the ciliary muscle contracts and the lens rounds up, while for distant vision the lens flattens out, so that the eye may be focused on distant objects Vitreous body Consists of colorless, transparent jelly-like substance in which there is a meshwork of fine fibrils, occupies the vitreous chamber, the space between lens and retina Helps maintain the shape of eyeball and supports the retina Refractive media of eye Bend entering light waves and focus them on the retina Cornea Aqueous humor Lens Vitreous body Accessory organs of eye Eyelids Conjunctiva Lacrimal apparatus Ocular muscles Connective tissue in the orbit Eyelids (from without inwards ) Skin, extremely thin Subcutaneous areola tissue, loose and delicate Muscular layer: orbicularis oculi Tarsus, formed by dense connective tissue in which the tarsal glands embedded Lined by palpebral conjunctiva Function: to protect, open, and close eye Tarsus Conjunctiva Three parts Palpebral conjunctiva : lining inner surface of eyelids Bulbar conjunctiva : lining anterior part of sclera, up to corneal margin Conjuntival fornix (superior and inferior): line of reflection of bulbar and palpebral conjunctiva Conjunctival sac Lacrimal apparatus Lacrimal gland Oval 2-cm, occupies fossa for lacrimal gland Ducts (6~10 in number): empty into anterior region of superior fornix of conjunctiva Secrets tears, which move across eyeball to medial angle,protect and moisten eye Lacrimal passages Lacrimal puncta opening to lacrimal ductules, one on each eylid margin near medial angle Lacrimal ductules : one in each lid, pass medially, join and enter lacrimal sac Lacrimal sac within fossa for lacrimal sac, opening into nasolacrimal duct Nasolacrimal duct courses 2 cm inferiorly and opens into inferior nasal meatus Tear is produced by lacrimal gland Passes through superior conjunctival fornix into conjunctival sac Then it is drained through lacrimal punctum, lacrimal ductule, lacrimal sac and nasolacrimal duct into inferior nasal meatus. Ocular muscles Muscle Action Nerve supply levator palpebrae superioris Raises upper eyelid Ⅲ Superior rectus Turns eyeball superomedially Ⅲ Inferior rectus Turns eyeball inferomedially Ⅲ Medial rectus Turns eyeball medially Ⅲ Lateral retus Turns eyeball laterally Ⅵ Superior obliquus Turns eyeball inferolaterally Ⅳ Inferior obliquus Turns eyeball superolaterally Ⅲ Ocular muscles 眼球外肌 Connective tissue in the orbit Sheath of eyeball : a thin membrane, which surrounds the eyeball from optic nerve to corneoscleral junction, permits the eyeball to move in the orbit without friction Adipose body of orbit : lies between sheath of eyeball and the orbit acts as a protective cushion and shock sorber for the eyeball Vessels of eye Ophthalmic artery Branch of internal artery Branches-central artery of retina Enters optic nerve, passes toward the optic disk and then fans out to supply the retina Four branches: superior and inferior nasal or temporal arteriole of retina Ophthalmic vein Superior ophthalmic vein communicates with facial vein anteriorly, exits posteriorly via superior orbital fissure to drain into cavernous sinus Inferior ophthalmic vein lies on floor of orbit and communicates with pterygoid plexus, exits via superior orbital fissure to drain into cavernous sinus The Vestibulocochlear Organ Liu jiao Binzhou Medical University Department Of Anatomy General features Three parts External ear : collects sound waves Middle ear : transmits sound waves Internal ear : contains the vestibulocochlear organ concerned with equilibration and hearing External ear Auricle External acoustic meatus Tympanic membrane Auricle External acoustic meatus A slender canal that extends from external acoustic pore to tympanic membrane Two parts Cartilaginous part- lateral 1/3 Bony part-medial 2/3 Lined by a layer of thin skin. This S-shaped passage medially, at first forward and upward, then backward and, finally forward and downward. Tympanic membrane A thin oval membrane Two parts Flaccid part : upper 1/4 Tense part : lower 3/4 Umbo of tympanic membrane Cone of light Middle ear Tympanic cavity Auditory tube Mastoid antrum and mastoid cells Tympanic cavity An air-containing cavity locates within petrous portion of temporal bone Walls Roof Medial wall lateral wall Posterior wall Anterior wall Floor Walls Roof or tegmental wall formed by tegmen tympani, separates tympanic cavity from middle cranial fossa Floor or jugular wall separates the cavity from superior bulb of internal jugular vein Anterior wall or carotid wall separates tympanic cavity from carotid canal, superiorly lies two openings: Upper opening for tensor tympani muscle Lower opening for auditory tube, which communicates with nasopharynx Posterior wall or mastoid wall Aditus of mastoid antrum Pyramid lateral wall or membranous wall -tympanic membrane with epitympanic recess superiorly Medial wall or labyrinthine wall Promontory Fenestra vestibuli Fenestra cochleae covered by secondary tympanic membrane Prominence of facial canal Acute otitis media Perforation, inflammation or trauma Auditory ossicles Consists of chain of three bones: Malleus Incus Stapes Articulate by synovial joints Transmit vibration of tympanic membrane to footplate of stapes in fenestra vestibule. Muscles of auditory ossicles Tensor tympani Stapedius Auditory tube About 3~4 cm long, extends from nasopharynx posteriorly, laterally, and upward to tympanic cavity Two parts Bony part: posterolateral 1/3 Cartilaginous part: medial 2/3 Functions to equalize air pressure on either side of tympanic membrane In childhood, it is shorter, wider and more horizontal than in adult Mastoid antrum and mastoid cells Mastoid antrum: a small chamber between tympanic cavity and mastoid cells Mastoid cells: contain a group of air cells within mastoid process of temporal bone Internal ear General features Lies within the petrous portion of temporal bone Key contents of internal ear Bony labyrinth contains perilymph Membranous labyrinth is filled with endolymph and contains the sensory organs Bony labyrinth Cochlea Vestibule Bony semicircular canals Cochlea It somewhat resembles a snail’s shell Consists of Modiolus Cochlear spiral canal makes two and one-half spinal turns around the modiolus Osseous spiral lamina Scala vestibuli Scala tympani Vestibule Hollow bony space Contains utricle and saccule Bony semicircular canals (anterior, posterior, and lateral) posteriorly Semicircular duct in each Canal at right angles to each other Dilated ampulla in each canal,called bony ampullar Membranous labyrinth Cochlear duct Utricle and saccule Semicircular ducts Cochlear duct Contains spinal organ (of Corti), the sound receptors lies on tympanic wall of cochlear duct Utricle and saccule Contain macular utricli and macular sacculi, end organs of balance, which respond to linear acceleration and deceleration, static of gravity Semicircular ducts Each duct has a membranous ampullae Containing crista ampullaris, receptors of balance that respond to rotational acceleration in three different planes Conduction of sound Sound waves Cochlear nerve Air-conduction of sound Bony- conduction of sound Sound waves Skull Bony labyrinth Endolymph within cochlear duct Cochlear nerve Perilymph Spinal organ Internal acoustic meatus Extends from internal acoustic pore to fundus of internal acoustic meatus Facial, vestibulocochlear nerves and vessles of labyrinth pass through the fundus of internal acoustic meatus Thank you