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EVOLUTION/POPULATION GENETICS CH. 15-16 SOME INFLUENCES ON DARWIN’S THOUGHT • • • • • LAMARCK MALTHUS LYELL HUTTON WALLACE : Contributor’s to Darwin’s thinking included: • Charles Lyell – uniformatarianism • Georges Cuvier – species extinction (Catastrophism) • Thomas Malthus – struggle for existence (resources) : Contributor’s to Darwin’s thinking included: • James Hutton - Gradualism • John Baptiste Lamarck – Inheritance of acquired Characteristics and Law of Use and Disuse • Alfred Russel Wallace – organisms evolved from common ancestors Lamarck’s Theory of Evolution • Jean-Baptiste Lamarck, 1809 • One Of First Scientists To Understand That Change Occurs Over Time • Stated that Changes Are Adaptations To Environment acquired in an organism’s lifetime • Said acquired changes were passed to offspring Lamarck’s Theory of Evolution • Idea called Law of Use and Disuse • If a body part were used, it got stronger • If body part NOT used, it deteriorated Lamarck’s Theory of Evolution • Inheritance Of Acquired Traits – Traits Acquired During Ones Lifetime Would Be Passed To Offspring Clipped ears of dogs could be passed to offspring! Lamarck’s Mistakes • Lamarck Did NOT Know how traits were inherited (Traits are passed through genes) • Genes Are NOT Changed By Activities In Life • Change Through Mutation Occurs Before An Organism Is Born Population Growth • Thomas Malthus, 1798 • Economist • Observed Babies Being Born Faster Than People Were Dying • Population size limited by resources such as the Food Supply The Struggle for Existence • Malthus’ Influence: – High Birth Rates & Limited Resources Would Force Life & Death Competition • Each Species Struggles For: – Food – Living Space – Mates Population Growth • Malthus Reasoned That If The Human Population Continued To Grow Unchecked, Sooner or Later There Would Be Insufficient Living Space & Food For Everyone • Death Rate Will Increase To Balance Population size & Food Supply Population Growth • Darwin Realized Malthus’s Principles Were Visible In Nature • Plants & Animals Produce Far More Offspring Than Can Be Supported – Most Die – If They Didn’t – Earth Would Be Overrun Charles Lyell • Proposed theory of Uniformitarianism • Geological processes at uniform rates building & wearing down Earth’s crust • Proposed that the Earth was millions of years instead of a few thousand years old Principles of Geology • Published by Lyell Just Before The Beagle Set Sail & read by Darwin • Explained Geological Processes That Shaped The Earth • Helped Darwin Understand Sea Shells In The Andes Mountains At 12,000+ Feet – Expanded Earth’s Age Hutton’s Theory of Geological Change •James Hutton, 1795, Scottish geologist •Studied invertebrate fossils in Paris Museum •Described The Geological Forces That Have Changed Life on Earth Over Millions of Years (erosion, earthquakes, volcanoes…) Hutton’s Theory of Geological Change • Changes in Earth’s crust due to slow continuous processes • Idea Known as Gradualism Wallace’s Contribution • Alfred Russel Wallace Independently came to same Conclusion as Darwin that species changed over time because of their struggle for existence • When Darwin read Wallace’s essay, he knew he had to publish his findings Darwin's Theory 1. Individual Organisms In Nature Differ From One Another. Some Of This Variation Is Inherited 2. Organisms In Nature Produce More Offspring Than Can Survive, And Many Of These Offspring Do No Reproduce Darwin's Theory 3. Because More Organisms Are Produced Than Can Survive, Members Of Each Species Must Compete For Limited Resources 4. Because Each Organism Is Unique, Each Has Different Advantages & Disadvantages In The Struggle For Existence Darwin's Theory 5. Individuals Best Suited To Their Environment Survive & Reproduce Successfully – Passing Their Traits To Their Offspring. 6. Species Change Over Time. Over Long Periods, Natural Selection Causes Changes That May Eventually Lead To New Species Darwin's Theory 7. Species Alive Today Have Descended With Modifications From Species That Lived In The Past 8. All Organisms On Earth Are United Into A Single Tree Of Life By Common Descent Section 15-3 Concept Map Evidence of Evolution includes The fossil record Geographic distribution of living species Homologous body structures Similarities in early development which is composed of which indicates which implies which implies Physical remains of organisms Common ancestral species Similar genes Similar genes Homologous Structures Section 15-3 Turtle HOMOLOGOUS STRUCTURES Alligator Ancient lobe-finned fish Bird Mammal Homologous Body Structures • Not All Serve Important Functions – Vestigial Organs • Appendix In Man • Legs On Skinks Evidence for Evolution - Comparative Embryology Similarities In Embryonic Development Fossil Record • Earth is Billions of Years Old • Fossils In Different Layers of Rock (sedimentary Rock Strata) Showed Evidence Of Gradual Change Over Time Geographic Distribution of Living Species Section 15-3 Beaver Beaver NORTH AMERICA Muskrat Muskrat Beaver and Muskrat Coypu Capybara Capybara SOUTH AMERICA Coypu Coypu and Capybara Similarities in DNA Sequence Evolutionary Time Scales Macroevolution: Long time scale events that create and destroy species. Evolutionary Time Scales Microevolution: Short time scale events (generationto-generation) that change the genotypes and phenotypes of populations VARIATION & GENE POOLS: GENE POOL, Section 16-1 RELATIVE FREQUENCY OF GENES/ALLELES Sample Population 48% heterozygous black Frequency of Alleles allele for brown fur allele for black fur 16% homozygous black 36% homozygous brown Relative frequency # of times an allele occurs in a gene pool, compared w/ the # of times other alleles for the same gene occur MUTATION • THE ULTIMATE SOURCE OF GENETIC VARIATION!!!!!! Phenotypes for Single-Gene Trait: The # of phenotypes a given Section 16-1 trait has is determined by how many genes control the trait Frequency of Phenotype (%) 100 Controlled by a single gene that has 2 alleles leads to 2 distinct phenotypes 80 60 40 20 0 Widow’s peak Phenotype No widow’s peak Controlled by 2/ more genes (2/ more alleles Frequency of Phenotype Generic Bell Curve for Polygenic Trait: many Section 16-1 possible genotypes and phenotypes Phenotype (height) NATURAL SELECTION ON POLYGENIC TRAITS: 3 MODELS Section 16-2 Key Directional Selection Low mortality, high fitness Food becomes scarce. High mortality, low fitness Favors traits at 1 extreme of a range of traits Graph of Stabilizing Selection Section 16-2 Stabilizing Selection Individuals with the most common trait are most adapted, while individuals who differ from the norm are poorly adapted. Key Low mortality, high fitness High mortality, low fitness Birth Weight Selection against both extremes keep curve narrow and in same place. Graph of Disruptive Selection Section 16-2 Disruptive Selection Low mortality, high fitness High mortality, low fitness Population splits into two subgroups specializing in different seeds. Beak Size Number of Birds in Population Key Number of Birds in Population Largest and smallest seeds become more common. Beak Size When both extreme phenotypes are favored by natural selection Balancing Selection Natural selection acts in opposite directions *** “Heterozygote Advantage”- ind who is heterozygous for a particular gene has a greater fitness than a homozygous ind EX: Distribution of sickle-cell allele coincides with the occurrence of malaria SS Normal hemoglobin ss Sicke-cell disease Ss- codominance (protects against malaria) GENTIC DRIFT: A RANDOM CHANGE IN ALLELE FREQUENCY • IN SMALL POPULATIONS, INDIVIDUALS THAT CARRY A PARTICULAR ALLELE MAY LEAVE MORE DESCENDANTS THAN OTHER INDIVIDUALS, JUST BY CHANCE. OVER TIME, A SERIES OF CHANCE OCCURRENCES OF THIS TYPE CAN CAUSE AN ALLELE TO BECOME COMMON IN A POPULATION. • FOUNDER EFFECT: A SITUATION IN WHICH ALLELE FREQUENCIES CHANGE AS A RESULT OF THE MIGRATION OF A SMALL SUBGROUP OF A POPULATION. Section 16-2 Genetic Drift Sample of Original Population In small populations, Descendants Founding an Population A can become allele more or less common by chance (explain how allele frequencies can fluctuate Founding Population B unpredictably from 1 gen to the next) Section 16-2 Genetic Drift Sample of Original Population Founding Population A Founding Population B Descendants FOUNDER EFFECT: 2 small groups from a large, diverse population could produce new populations that differ from the original population Section 16-2 Genetic Drift Sample of Original Population Descendants Founding Population A Founding Population B EVOLUTION VERSUS GENETIC EQUILIBRIUM HARDY-WEINBERG PRINCIPLE: ALLELE FREQUENCIES IN A POPULATION WILL REMAIN CONSTANT AS LONG AS FIVE CONDITIONS (FACTORS) REMAIN CONSTANT. GENETIC EQUILIBRIUM IS REACHED. (IS THE POPULATION EVOLVING?) 5 CONDITIONS REQUIRED TO MAINTAIN GENETIC EQUIIBRIUM 1. 2. 3. 4. 5. RANDOM MATING - Select mates w/o bias LARGE POPULATION - Genetic drift does not affect large pops NO MOVEMENT INTO OR OUT OF THE POPULATION - no intro of new allele NO MUTATION - no new allele introduced NO NATURAL SELECTION - No phenotype can have selective advantage DOES THIS EVER HAPPEN????? • In 1908, Hardy and Weinberg independently demonstrated that DOMINANT ALLELES DO NOT REPLACE RECESSIVE ALLELES IN A POPULATION! HARDY-WEINBERG EQUATION You have sampled a population in which you know that the percentage of the homozygous recessive genotype (aa) is 36%. Using that 36%, calculate the following: The frequency of the "aa" genotype. The frequency of the "a" allele. The frequency of the "A" allele. The frequencies of the genotypes "AA" and "Aa." The frequencies of the two possible phenotypes if "A" is completely dominant over "a." • FREQUENCY OF aa GENOTYPE: 36% (GIVEN) • FREQUENCY OF a ALLELE:The frequency of aa is 36%, which means that q2 = 0.36, by definition. If q2 = 0.36, then q = 0.6, again by definition. Since q equals the frequency of the a allele, then the frequency is 60%. • The frequency of the "A" allele. Answer: Since q = 0.6, and p + q = 1, then p = 0.4; the frequency of A is by definition equal to p, so the answer is 40%. • The frequencies of the genotypes "AA" and "Aa." Answer: The frequency of AA is equal to p2, and the frequency of Aa is equal to 2pq. So, using the information above, the frequency of AA is 16% (i.e. p2 is 0.4 x 0.4 = 0.16) and Aa is 48% (2pq = 2 x 0.4 x 0.6 = 0.48). • The frequencies of the two possible phenotypes if "A" is completely dominant over "a." Answers: Because "A" is totally dominate over "a", the dominant phenotype will show if either the homozygous "AA" or heterozygous "Aa" genotypes occur. The recessive phenotype is controlled by the homozygous aa genotype. Therefore, the frequency of the dominant phenotype equals the sum of the frequencies of AA and Aa, and the recessive phenotype is simply the frequency of aa. Therefore, the dominant frequency is 64% and, in the first part of this question above, you have already shown that the recessive frequency is 36%. AS NEW SPECIES EVOLVE (SPECIATION), POPULATIONS BECOME REPRODUCTIVELY ISOLATED FROM EACH OTHER Section 16-3 Reproductive Isolation results from Isolating mechanisms which include Behavioral isolation Geographic isolation Temporal isolation produced by produced by produced by Behavioral differences Physical separation Different mating times which result in Independently evolving populations which result in Formation of new species . Flowchart Section 17-4 Species that are Unrelated form Related in under under in in Interrelationshiops Similar environments Intense environmental pressure Small populations Different environments can undergo can undergo can undergo can undergo can undergo Coevolution Convergent evolution Extinction Punctuated equilibrium Adaptive radiation a. Divergent evolution when 2/ more species originate from a common ancestor Ex: flipper of whale and limb of human b. Convergent evolution process by which unrelated organisms come to resemble each other Ex: torpedo shape of shark and penguin/ wing of insect and bat c. Coevolution process by which 2 species evolve in response to changes in each other over time