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Introduction to Animal Diversity Packet #76 Chapter #32 Animal Diversity Biologists have identified 1.3 million living species if animals. Estimates put the range much higher 10 – 200 million Evolutionists believe that the chanoflagellates, a colonial flagellated protist, was the start of Kingdom Animalia. Characteristics Most Common to Animals Eukaryotic Multicellular Heterotrophic Animals use enzymes to digest their food after they have ingested it. Stores energy as fat (long term) or as glycogen (short term) Reproduction in Animals Reproduction—The Obvious Most animals reproduce sexually and have a diploid stage that is dominant in the life cycle. Sperm and egg unite to form a zygote Zygote undergoes cleavage Multiple cell divisions result in the development of a hollow ball of cells Blastula Blastula undergoes gastrulation. Embryonic tissues are formed Developmental stage is called the gastrula. Reproduction—The Sometimes Not So Obvious Some animals develop directly into adults After transient stages of maturation However, life cycle of many animals include larval stages. Larva Sexually immature form of an adult Morphologically distinct Usually eats different food Inhabits different areas than the adult Must undergo metamorphosis to become an adult Developmental Genes Across Kingdom Animalia Animals share a unique homeobox Family of genes Hox genes The number of hox genes is correlated with the complexity of the animal’s anatomy. Categorizing Kingdom Animalia Animals are classified on major features of animal body plans. Symmetry Tissue Complexity Cephalization Body Cavities Body Plans Symmetry Introduction There are two types of symmetry Radial Symmetry Bilateral Symmetry Symmetry Radial Symmetry Describes how the parts of an animal radiate from the center. Any imaginary slice through the central axis divides the animal into mirror images. Sea anemones have a top (oral, mouth) side and a bottom (aboral) side. Symmetry II Bilateral Symmetry Describes a two sided body plan. Animal has a left side and right side Imaginary slice can only be placed in one location in order to divide the animal mirror images. Lobster has a dorsal (top) side, a ventral (bottom) side, a left and right side, an anterior (head) with a mouth and a posterior (tail) end. Symmetry III Animals can be categorized according to the symmetry of their bodies or lack of it. Symmetry Reflects Lifestyle Radial animals are sessile or planktonic Bilaterial animals more actively from one place to another The nervous system enables these organisms to move. Tissue Tissue I As a young embryo develops, embryonic tissue, called germ layers, are produced via gastrulation. There are three germ layers Ectoderm Endoderm Mesoderm Tissue II Ectoderm Outer layer Gives rise to the body covering and the nervous system Endoderm Inner layer Gives rise to the lining of the gut (archenteron) and other digestive organs Mesoderm Middle layer Gives rise to most other body structures. Including muscle Diploblastic vs. Triploblastic Diploblastic Animals with only two layers Ectoderm and Endoderm Jellies Corals Comb jellies Triploblastic Animals with all three layers Include all bilaterally symmetric animals. Triploblastic Animals & Body Cavities Functions of Body Cavities Provides protection to internal organs Allow organs to grow and move independently of the outer body wall. Body Cavity I Triploblastic animals have traditionally been classified as Acoelomates [ey-see-luh-meyt] No body cavity Lack a coelom. [see-luhm] Pseudocoelomate [soo-dohsee-luh-meyt, -si-loh-mit] Body cavity not completely lined with mesoderm Body cavity formed from the blastocoel. Coelomate True coelom Body cavity completely lined with mesoderm. Cushions the internal organs and protects them. Formation of the Coelom [see-luhm] Coeloms can be divided into two categories based on how it is developed. During gastrulation, developing digestive tube forms the archenteron. Protostomes Development of the coelom forms from splits in the mesoderm Schizocoelous Development Deuterostomes Development of the coelom forms from outpocketing of the mesodermal tissue of the archenteron. Enterocoelous Development The Coelomates Introduction I Protostomia Mollusks Annelids Arthropods Deuterostomia Enchinoderms Chordates Protostomes Blastopore develops into the mouth Undergo spiral and determinate cleavage Spiral cleavage Describes how the planes of cell division are diagonal to the vertical axis of the embryo. Smaller cells lie in the grooves between larger, underlying cells Determinate cleavage Indicates that the developmental fate of each embryonic cell is determined at fertilization. If cell is isolated it will form an inviable embryo. Deuterostomes Blastopore typically becomes the anus. Undergo radial and indeterminate cleavage. Radial cleavage Cleavage planes are either parallel or perpendicular to the vertical axis of the egg Indeterminate cleavage Each cell produced by early cleavage divisions has the capacity to develop into a complete embryo. Protostomes vs. Deuterostomes Review Symmetry Symmetry Radial Symmetry Bilateral Symmetry Sea anemones Lobster Tissue Germ Layers Ectoderm Mesoderm Endoderm Outer layer Middle layer Inner Layer Body Covering Nervous System Muscle Other Body Structures Lining of Gut Digestive System Germ Layers Body Cavity Body Cavity Diploblastic Triploblastic Acoelomates Psuedocoelomates Protostomes Coelomates Deuterostomes Body Cavity Coelomates Protostomes Blastopore Mouth Spiral Cleavage Deuterostomes Determinate Cleavage Blastopore Anus Radial Cleavage Indeterminate Cleavage