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Section 9.2 (Part 1) SBI 3U1 pp. 360 - 365 SPECIATION: HOW SPECIES FORM MICROEVOLUTION Changes in the gene pool of a population over time which result in relatively small changes to the organisms in the population MACROEVOLUTION Changes in organisms which are significant enough that, over time, the newer organisms would be considered an entirely new species. The new organisms would be unable to mate with their ancestors, assuming we were able to bring them together. SPECIES Species - A group of actually or potentially interbreeding natural populations that ordinarily do not interbreed with other such groups even when there is opportunity to do so. SPECIATION Speciation The formation of new species from existing species. The formation of new species is also sometimes called macroevolution TYPES OF SPECIATION Allopatric Speciation The formation of two or more species often requires geographical isolation of subpopulations of the species. Only then can natural selection or perhaps genetic drift produce distinctive gene pools. Sympatric Speciation Species diverge while inhabiting the same space and become reproductively isolated ALLOPATRIC SPECIATION Allopatric speciation by migration Removal of the geographic barrier Lets try this again SYMPATRIC SPECIATION What might keep two subpopulations from interbreeding when reunited geographically? There are several isolating mechanisms. REPRODUCTIVE ISOLATION ISOLATING MECHANISMS Pre-zygotic – Before the zygote is formed mating is prevented Post-zygotic – After the zygote is formed mating is not prevented, but the hybrids have reduced fitness (reduced chance of survival) PRE-ZYGOTIC ISOLATING MECHANISMS Pre-zygotic - Factors which cause species to mate with their own kind Temporal isolation - Individuals of different species do not mate because they are active at different times of day or in different seasons. E.g. orchid & blooming time Ecological isolation - Individuals mate in their preferred habitat, and therefore do not meet individuals of other species with different ecological preferences. E.g. Garter snake (meadow vs. near water) Behavioural isolation - Potential mates meet, but choose members of their own species. E.g. Songs of birds Mechanical isolation - Copulation is attempted, but transfer of sperm does not take place. E.g. Insects Gametic Isolation - Sperm transfer takes place, but egg is not fertilized. POST ZYGOTIC ISOLATING MECHANISMS Post-zygotic. Genomic incompatibility, hybrid inviability or sterility. Zygotic mortality/Hybrid Breakdown – Hybrid is viable and fertile, but hybrid’s offspring is sterile or weak. E.g. Cotton plant hybrid has offspring that die as seeds or early in development. Hybrid Inviability - Hybrid embryo forms, dies in early development before birth. E.g. Hybrid embryo between sheep and goat. Hybrid sterility - Hybrid is viable, but resulting adult is sterile. E.g. Mule from female horse and male donkey p. 360, Figure 9.12 TO DO LIST Section 9.2 p. 363 #13 – 18, p. 365 #19 – 24 Unit Test for Evolution Monday Nov 23, 2015