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1.Compare and contrast the protostomes and deuterostomes. Protosomes Blastopore becomes mouth & 2 becomes anus Deuterosomes nd opening Blastopore becomes anus & 2nd opening becomes mouth Coelom formed via schizocoely Coelom formed via enterocoely spiral, determinate cleavage at 4 —> 8 cell division radial, indeterminate cleavage at 4 —> 8 cell division circulatory system primitively dorsal and nervous system primitively ventral circulatory system primitively ventral and circulatory system primitively ventral Phyla Mollusca, Annelida, Arthropoda and others Phyla Echinodermata, Chordata and other Deuterostome Protostome Future intestine Blastopore Mouth Hemichordata, Future Anus Future mouth Future intestine Blastopore 2 Radial cleavage Gut Blastocoel Blastocoel Mesoderm Pocket of gut Coelom Mesoderm 4 Regulative embryo 4-cell embryo 1 blastomere excised 4-cell embryo 1 blastomere excised 2 normal larvae 2. Describe the enterocoel hypothesis of coelom origin and discuss the implications. Enterocoelous development is the stage of embryological development of deuterostomes in which the coelom forms. The stage starts with the gastrula; as the archenteron forms, pockets of migrating cells also form, creating another layer between the endoderm and ectoderm, the mesoderm. These pockets gradually expand to form the coelom. Enterocelous development is advance in comparison to schizocoelous development due to the formation of the archenterons. 3. Why was the development of a true coelom such an important step in the evolution of higher animals? A true coelom is a fluid-filled body cavity formed from mesodermal tissue. It lies between the outer body wall of epidermal tissue and the gut or digestive tract. Evolutionary advantages of coelom can be outlined as follows : 1. The coelomate body plan is a “tube within a tube”. Because this tube is filled with fluid, it allows fluid circulation, even in primitive animals that lack circulatory systems. 2. Fluids (like water) are relatively incompressible. The fluid-filled coelom can therefore act as a type of rigid skeleton, or hydrostatic skeleton. The muscles now have something solid to push against. 3. The coelom allows for an open digestive tract, with a mouth at one end, an anus at the other, and this tract can be increased by coiling within the coelom so that it is many times longer than the animal itself. 4. Animals like flatworms, on the other hand, with one opening into a hollow cavity, are limited in how fast they can eat, digest, and excrete. 5. A coelom allows digestion independent of movement. Gut action need no longer depend on the muscular contractions generated by the animals movements. 6. There is more space for the internal organs to develop, especially the gonads, and large numbers of eggs and sperm can be stored in the coelom as well. 7. And finally, the combination of a coelom and bilateral symmetry opens up an entirely new evolutionary pathway, in which parts of the body can be adapted to perform special functions. This new pathway, which has ultimately given insects dominion over all other life, is called segmentation. 4. What are the major characteristics of the phylum Cnidaria? 1. Multicellular, metazoan animals which are radially symmetrical about an oral aboral axis. 2. 3. 4. Two types of body forms, medusae and polyps. The animals have reached the tissue level of organisation. The bodies are derived from two embryonic cell layers (diploblastic), the ectoderm and endoderm which are separated by a third gelatinous layer called the mesoglea (this is not the same as “Mesoderm”). The main body cavity, the gastrovascular cavity has a single opening that serves as both the mouth and anus. 5. 6. 7. 8. 9. 10. Support of the organisms is generated either through a calcareous exoskeleton or through a rudimentary hydrostatic skeleton. The inner, outer or both surfaces are possess special stinging cells, cnidocytes. Locomotion and co-ordination are generated by epithelio-muscular cells and a primitive nerve net. The animals have no excretory, circulatory or respiratory systems. Reproduction is by asexual budding or sexual reproduction. 5. What is a possible consequence of a one-way digestive tract? Cellular specialization in one way digestive tract allows ingested food to be processed along a one way passage, with unassimilated matter being ejected out the anus. It does not return and passed back out the oral opening as observed in cnidarians and most platyhelminthes 6. What criteria are used to divide the animal kingdom into phyla? Phylum is the second highest taxonomic classification for the kingdom Animalia. It lies in between kingdom level and class level. Each kingdom is subdivided into Phylums which are in turn differentiated into classes of organism. Phylum category is recognized on the basis of a fundamental body plan and secondly organisms of one phyla evolved from a common ancestry. 7. Compare and contrast the degree to which members of the phyla Porifera, Cnidaria and Platyhelminthes exhibit nervous control of body functions. Porifera Nervous system is absent Cnidaria Nervous system organized as a “nerve net”, but lacking a brain. Platyhelminthes Nervous system consists of : a) “brain” and sensory organs located on anterior portion (cephalization) b) “eyespots” are sensitive to the presence of light c) lateral nerve cords connected by transverse nerves forming a “nerve ladder”