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Individual organisms differ, and some of this variation is heritable Organisms produce more offspring than can survive, and many that do survive do not reproduce Because more organisms are produced than can survive, they compete for limited resources Each unique organism has different advantages and disadvantages in the struggle for existence. Individuals best suited to their environment survive and reproduce most successfully. These organisms pass their heritable traits to their offspring. Other individuals die or leave fewer offspring. This process of natural selection causes species to change over time. Species alive today are descended with modification from ancestral species that lived in the distant past. This process, by which diverse species evolved from common ancestors, unites all organisms on Earth into a single tree of life. 1. Mutations 2. Genetic shuffling that results from sexual reproduction *The number of phenotypes produced for a given trait depends on how many genes control the trait 1. single-gene trait = leads to 2 phenotypes, ex: widow’s peak, tongue rolling 2. polygenic trait = many possible genotypes and phenotypes, ex: height Gene pool = consists of all genes, including all the different alleles, that are present in a population Relative frequency = number of times that the allele occurs in a gene pool compared with the number of times other alleles for the same gene occur. (%) Evolution = any change in the relative frequency of alleles in a population ◦ If the relative frequency of an allele changes over time, that means the population is evolving A. Fossil record ◦ Derived Traits = newly evolved features, such as feathers, that do not appear in the fossils of common ancestors ◦ Ancestral Traits = primitive features, such as teeth and tails, that do appear in ancestral forms B. Comparative Anatomy ◦ Homologous structures = anatomically similar structures inherited from a common ancestor Arm, foreleg, fin, wing ◦ Vestigial structures = structures that are reduced forms of functional structures in other organisms snake pelvis, kiwi wings, human appendix ◦ Analogous structures = can be used for the same purpose and can be superficially similar in construction but are not inherited from a common ancestor Bird wings and insect wings Adaptation = a trait shaped by natural selection that increases an organism’s reproductive success Fitness = measure of the relative contribution an individual trait makes to the next generation Types of Adaptation Camouflage = morphological adaptations that allow them to blend in with their envs. Mimicry = one species evolves to resemble another species (kingsnake and coral snake) Antimicrobial resistance Hardy-Weinberg principle = states that when allele frequencies in a population remain constant a population is in genetic equilibrium ◦ P2 + 2pq + q2 = 1 p = dominant, q = recessive ◦ Homozygous dom. + heterozygous + homozygous recessive = 1 Genetic equilibrium = the situation in which allele frequencies remain constant 5 conditions are required to maintain genetic equilibrium from generation to generation ◦ ◦ ◦ ◦ ◦ 1. 2. 3. 4. 5. Random mating Large population No movement into or out of the population (No interbreeding) No mutations No natural selection A. Natural selection = process by which favorable traits that are heritable become more common in successive generations of a population of reproducing organisms ◦ individuals with favorable phenotypes are more likely to survive and reproduce than those with less favorable phenotypes 1. Individuals at one end of the curve have higher fitness than individuals in the middle or at the other end ◦ Ex. Small and medium-sized seeds start to run low and only large seeds finches with large, thick beaks survive and reproduce so average beak size would increase 2. Individuals near the center of the curve have higher fitness than individuals at either end of the curve ◦ Ex. Babies that are too small or too large are less likely to survive (average is selected for) 3. Individuals at the upper and lower ends of the curve have higher fitness than individuals near the middle ◦ Ex. Snakes live at edge of forest. Light and dark are selected for. Medium color is disadvantage Natural selection in which change in frequency of a trait is based on the ability to attract a mate ◦ ex. Brighter and bigger tails in peacocks ◦ Larger size and antlers in deer Random change in allele frequency due to chance ◦ More extreme in small populations ◦ May occur when a small group of individuals colonizes a new habitat A situation in which allele frequencies change as a result of the migration of a small subgroup of a population Occurs when a population declines to a very low number and then rebounds Speciation = formation of new species ◦ A population must diverge and then be reproductively isolated Reproductive isolation = when the members of 2 pops. cannot interbreed and produce fertile offspring A physical barrier divides one population into 2 or more populations A species evolves into a new species without a physical barrier Prezygotic isolating mechanisms = operate before fertilization occurs ◦ Keep species separate Postzygotic isolating mechanisms = operate after fertilization ◦ hybrid remains infertile (liger, mule) 1. Behavioral isolation = occurs when 2 pops. are capable of interbreeding but have differences in courtship rituals or other reproductive strategies that involve behavior ◦ Ex. Eastern and western meadowlarks have overlapping ranges but do not mate with each other because they have diff. mating songs 2. Geographic isolation = 2 pops. are separated by geographic barriers such as rivers, mountains, or bodies of water ◦ Ex. Colorado river split pop. of Abert squirrels and 2 separate gene pools were formed to form a new species 3. Temporal isolation = 2 or more species reproduce at different times ◦ Ex. 2 pops. of orchids release pollen on different days so they cannot mate with each other, eventually they may become separate species Pg. 439 Explain Explain Explain Explain Adaptive radiation Coevolution Convergent evolution Rate of Speciation ◦ Gradualism ◦ Punctuated equilibrium