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The Struggle For Existence The Revolutionary idea of Evolution in determining species diversification History of Evolution Lamarck published a theory of evolution in 1809 Lamarck thought that species arose continually from nonliving sources. These species were initially very primitive, but increased in complexity over time due to some inherent tendency. This type of evolution is called orthogenesis. Orthogenesis continued an organism's acclimation to the environment could be passed on to its offspring; species never go extinct for example, he thought proto-giraffes stretched their necks to reach higher twigs. This caused their offspring to be born with longer necks. This proposed mechanism of evolution is called the inheritance of acquired characteristics Who might like this idea for structuring society? Darwin Observed Variability exists within a given species Variant traits may be passed on/inherited In this "struggle for existence", variants that were slightly better suited to the environment would be more likely to survive This does *NOT* mean “Survival of the Fittest” Big Damn Digital Post It Note Darwin Never Said Survival of the Fittest Variation of Traits passed on to population Populations Evolve; Not Individuals A population is a collection of individuals which have some distribution of traits: Darwin’s Says “I have called this principle, by which each slight variation, if useful, is preserved, by the term Natural Selection It is not the strongest of the species that survives, nor the most intelligent, but the one most responsive to change Intelligence is based on how efficient a species became at doing the things they need to survive In the survival of individuals and species, during the constantly-recurring struggle for existence, we see a powerful and ever-acting form of selection.” One More I think it inevitably follows, that as new species in the course of time are formed through natural selection, others will become rarer and rarer, and finally extinct. The forms which stand in closest competition with those undergoing modification and improvement will naturally suffer most. Final Exam Questions Is Natural Selection inconsistent with the idea of the Mechanical Universe? (what would Descartes Say?) Darwin – Origin of Species -1859 Mechanism for Evolution is Natural Selection New variants continually arose within populations Darwin observes this but does not understand what mechanism causes this Numerous instances of selecting the variant (due to higher survival probability) slowly leads to species modification/diversification Favorable Variations Drive Evolution Darwin: If certain variants will be preserved over time over other variants then that population will change over time in their composition. This is evolution by natural selection. But Darwin does not understand how the variations of traits are generated in the first place Natural Selection Random pairings of Gene Pool/traits (Darwin does not know this) Each pairing has a different survival probability within some Ecosystem If ecosystem is stable the end results is a distribution of most probable values (slow vs fast rabbits) This statistical process takes a large number of generations to manifest itself Statistical types of selection In complex creatures, the variation of traits may be quite large make it much harder to discern the average set of traits from one individual creature. Selection as “Deficiency Correction” advancement of the mean The most common action of natural selection is to remove unfit variants as they arise (randomly) via mutation. In other words, natural selection usually prevents unfit genes from increasing in frequency in the gene pool. Advancement of the Mean is dangerous and erroneous concept Evolution is not an optimization process – there is never enough time for that – so why do we pretend that is natural optimization instead of natural selection? – – Supports the Social Darwinist Agenda that society can be engineered Takes nature out of statistical equilibrium if it can optimize to produce the Master Race Each Black Dot in each distribution represent individuals that die out before passing on their genes Stabilization: Extreme ends of the distribution are disfavored. Moves every individual closer to the average state. Directional: What you think is evolution – removes the deficient Disruptive: Individuals with average traits are disfavored. Better leads to species diversification. Mutations will affect largest number of individuals (i.e. average). Leads to development of new species and arguably is the process that happens on Earth Darwin Abused: Survival of the Fittest Social Engineering will now use this new scientific theory of evolution via adaptation as the scientific basis on which to engineer society. Hebert Spencer: This survival of the fittest, which I have here sought to express in mechanical terms, is that which Mr. Darwin has called "natural selection” or the preservation of favored races in the struggle for life The Aberrated View of Spencer Spencer quickly adapted Darwinian ideas to his own ethical theories. The concept of adaptation allowed him to claim that the rich and powerful were better adapted to the social and economic climate of the time, and the concept of natural selection allowed him to argue that it was natural, normal, and proper for the strong to thrive at the expense of the weak. After all, he claimed, that is exactly what occurs in nature every day. Social Darwinism Not only was survival of the fittest natural, but it was also morally correct. it was morally incorrect to assist those weaker than oneself, since that would be promoting the survival and possible reproduction of someone who was fundamentally unfit. The American eugenics movement was relatively popular between about 1910-1930, during which 24 states passed sterilization laws and Congress passed a law restricting immigration from certain areas deemed to be unfit Summary Evolution requires genetic variation Genetic variation is caused by gene mutation Mutation is continuously occurring within a species Natural selection operates on this variation These variations are largely random Most adaptable variations survive independent of their “Fitness”