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Tropical Rain Forest Tropical rain forests are hot and wet year-round and are home to more species than all other biomes combined. The leafy tops of tall trees – extending from 50 to 80 meters above the forest floor – form a dense covering called a canopy. In the shade below the canopy, a second layer of shorter trees and vines forms a dense understory. Organic matter that falls to the forest floor quickly decomposes and the nutrients are recycled. The soil in a tropical rain forest is thin and nutrient-poor. Canopy dense covering formed by tall trees Understory shorter trees and vines Tropical Savanna Tropical savannas are characterized by warm temperatures and seasonal rainfall. Receiving more seasonal rainfall than deserts, but less than tropical forests, tropical savannas are characterized by a cover of grasses. Savannas are spotted with isolated trees and small groves of trees and shrubs. Compact soils, fairly frequent fires, and the action of large animals such as rhinoceroses prevent some savanna areas from turning into forests. Tall, perennial grasses and drought-resistant trees dominate the savanna. Animal Adaptations in the African Savanna Migration During the rainy season, birds, insects, and both large and small mammals thrive in the savannah, but the rainy season only lasts 6 to 8 months. During the dry season, surface water from the rain is quickly absorbed into the ground because the soil is extremely porous. Competition for water during the dry season is intense. Consequently, most birds and many of the large mammals migrate during the dry season in search of water. Because drought conditions are sometimes localized, the migration may be just to another area within the savannah. When drought conditions exist for a long time and over a wide area, the animals may migrate to another biome until the rainy season begins again. Although elephants do migrate, they have a physical adaptation that allows them to access water that is not available to other animals. Baobab trees store water in their large trunks. The elephant's physical strength and anatomy allow it to tear open the trunk of the baobab tree and to suck the water from it. An adaptation used by small burrowing animals is to remain dormant during times of drought--much like bears do during the winter in other biomes Lightning During the dry season, lightning frequently ignites the brown, dry grasses that cover the savannah. Many of the animals have adapted to living with the fires. The ability to fly or to run fast enables most birds and large mammals to escape the flames. Some birds, such as the Fork-tailed Drongos, actually are attracted to the active fires. These birds feast on fleeing or flame-roasted insects. Although small burrowing animals may not be able to outrun the flames, they frequently survive the fire by digging beneath the soil and remaining there until the flames pass by them. Why do giraffes have such long necks? Competition for Food The “competing browsers hypothesis” was originally suggested by Charles Darwin and only challenged recently. It suggests that competitive pressure from smaller browsers, such as kudu, steenbok, and impala, drove the elongation of the neck so giraffes could reach nutrients competitors could not. This advantage is real – giraffes can and do feed up to 5 m, while most of their competitors, kudu, can only feed up to about 2 m (7 ft).[ There is also research suggesting that browsing competition below 2 m is intense, and giraffes feed more efficiently (gaining more leaf biomass per bite) higher in the canopy. However, scientists disagree about just how much time giraffes spend feeding at levels unreachable to other browsers. Although giraffes can feed as low as 0.5 m and as high as 6 m off the ground, it appears that they most often feed between 2 and 4 m (7–14 ft). However, elephants also routinely feed at heights up to 5 m (they knock down only a minority of the trees they feed on), and are likely competitors at these heights. Competition for food with other giraffes could also favor the evolution of tall necks. Competition for females The other main theory, the sexual selection hypothesis, proposes that the long necks evolved as a secondary sexual characteristic, giving males an advantage in "necking" contests to establish dominance and obtain access to sexually receptive females. In support of this theory, males have proportionally larger necks than females, and males with longer, bigger necks are more successful in dominance displays and courtship behavior. However, a major criticism of this theory is that it fails to adequately explain why female giraffes also have long necks. Temperate Grasslands Characterized by a rich mix of grasses and underlain by some of the world’s most fertile soils, temperate grasslands – such as plains and prairies – once covered vast areas of the Midwest and central United States. Since the development of the steel plow, however, most have been converted to agricultural fields. Periodic fires and heavy grazing by large herbivores maintain the characteristic plant community. Temperature Grasslands generally have warm to hot summers and cold winters with moderate, seasonal precipitation. Grassland animals Herbivores: •Mule deer •Pronghorn antelopes, •Rabbits •Prairie dogs •Bison Predators: •Coyotes •Badgers •Wolves •Owls •Hawks •Prairie chickens Insects: •Ants •Grasshoppers Reptiles: •snakes Desert All deserts are dry – a desert biome is defined as having annual precipitation of less than 25 cm. Beyond that, deserts vary greatly, depending on elevation and latitude. Many undergo extreme temperature changes during the course of a day, alternating between hot and cold. The organisms in this biome can tolerate the extreme conditions. Soils in the desert are rich in minerals, but poor in organic material. Tundra The Tundra is characterized by permafrost, a layer of permanently frozen subsoil. During the short, cool summer, the ground thaws to a depth of a few centimeters and becomes soggy and wet. In winter, the topsoil freezes again. This cycling of thawing and freezing, which rips and crushes plant roots, is one reason that tundra plants are small and stunted. Cold temperatures, low precipitation, high winds, the short growing season, and humus-poor soils also limit plant height. Permafrost A layer of soil that stays frozen all year The Calliergon giganteum has adapted well to its cold climate. When it is not growing, it stores nutrients so new leaves can be made quickly next spring. The more leaves the more they can photosynthesize. It is adapted to the incredibly strong winds found in the tundra by growing near to the ground. Because it can grow under water it is protected from the drying winds and cold, dry air of the frozen tundra. Its long life and slow growth are probably adaptations to the short growing season and the cold. Cushion plants - grow in a low, tight clump and look like a little cushion. Cushion plants are more common in the tundra where their growth habit helps protect them from the cold. Temperate Forest Temperate forests contain a mixture of deciduous and coniferous tress. These forests have cold winters that halt plant growth for several months. In autumn, the deciduous trees shed their leaves. In the spring, small plants burst out of the ground and flower. Summers are often warm. Soils of temperate forests are often rich in humus, a material formed from decaying leaves and other organic matter that makes soil fertile. This biome is also characterized by year-round precipitation. Deciduous trees trees that shed their leaves in the autumn Food web of the temperate forest Boreal Forest (Taiga) Along the northern edge of the temperate zone are dense evergreen forests of conifers. (Conifers are trees that produce seed-bearing cones which usually have leaves shaped like needles.) These biomes are called boreal forests, or taiga. Winters are bitterly cold, but summers are mild and short – but long enough to allow the ground to thaw. The word boreal comes from the Greek word for “north,” reflecting the fact that boreal forests occur mostly in the Northern hemisphere. This area is characterized by acidic, nutrient poor soils. Conifers trees that produce seed-bearing cones and usually have leaves shaped like needles. Partial food web showing consumers of the Taiga.