* Your assessment is very important for improving the workof artificial intelligence, which forms the content of this project
Download View/Open
Survey
Document related concepts
Transcript
Taxodium Native of Mexico and North America Having deciduous leaves Taxodium ascendens – the pond cypress T. distichum – the bald cypress T. mucronatum –the Montezuma yew cypress Taxodium mucronatum ‘Big tree of Tule’ 16m in diameter 5,000 years old (probably the oldest living thing in the world) Unlike other cypress, this is evergreen Does not possess a single dead twig Girth 58 meters Height 42 meters Position Family – Cupressaceae Subfamily – Taxodiaceae (Trees that are evergreen or deciduous) 10 members under subfamily Athrotaxis (3) Cryptomeria (1) Cunninghamia (2) Glyptostrobus (1) Metasequoia (1) Sciadopitys (1) Sequoia (1) Sequoiadendron (1) Taiwania(1) Taxodium (3) •Monoecious •Ovuliferous scales are free almost free when young •2-9 ovules on each ovuliferous scale •2-9microspoangiaor pollen sacs on each microsporophyl •Pollen grains are without wings •Pollen grains lack prothallial cells Ecology T. adnascens: The Pond Cypress occurs within the range of Bald Cypress, but only on the southeastern coastal plain from North Carolina to Louisiana. It occurs in still backwater rivers, ponds and swamps without silt-rich flood deposits. T. mucronatum: The Montezuma Cypress occurs from the Lower Rio Grande Valley south to the highlands of southern Mexico, and differs from the other two species in being substantially evergreen. t is a riparian tree, occurring on the banks of streams and rivers, not in swamps like the Bald and Pond Cypresses T. distichum: The most familiar species in the genus is the Bald Cypress, native to much of the south-eastern United States, from Delaware to Texas, especially Louisiana and inland up the Mississippi River to southern Indiana. It occurs mainly along rivers with silt-rich flood deposits. Morphology The persistent branches with axillary buds near the ends of the year’s growth Two types of branches The deciduous branches attached to the persistent shoots but without axillary buds Leaves are soft, short, narrow and pointed and show two types of on the shoots arrangements On the persistent shoots are spirally arranged; on the deciduous shoots are two ranked Winter buds are small , round with overlapping, sharp and pointed scales occurring near the tip of the shoot Bark is green when young, turns brown and scaly in older trees Leaves at first glance appear compound-in reality, they are simple. The needles are arranged in two ranks (that is, they occur in exactly two rows on each side of the stem)--this is called distichous (hence the epithet distichum). Cypress Knees Plants that grow in swamps or marshy places have knee like out growths from roots arising vertically above the soil for about 1-3.5m in known as ‘Cypress knees’ Knees are soft and spongy Perform same function as pneumatophores or breathing roots Lower part of the trunk shows extreme swellings and is profusely covered with adventitious roots Knees shows 6-8 xylem groups Normal roots are diarch Stem Anatomy Anatomy of 5-year old stem: pith and 1st wood growth ring (primary xylem indicated with arrow) annual growth rings growth ring, tracheid, ray TLS of Wood RLS of Wood Form: A large tree with a pyramid-shaped crown, cylindrical bole, fluted or buttressed base and often with knees Monoecious – male and female cones generally on different branches Flowers: Males in drooping, long panicles; females are sub globose, have peltate scales, and tend to occur near the end of branches. Fruit: Cones are composed of peltate scales forming a woody, brown sphere with rough surfaces, 3/4 to 1 inch in diameter; cones disintegrate into irregular seeds. Cone Characteristics •Cones appear near the tips of the branches of the same year and are found in the axil of the scale leaves •Female cone are irregularly placed on shoots and occur in groups of two or three •Male cones are long drooping panicles •Female cones are ball shaped Male cone Male cone is small, round gradually turns to oblong on maturity Colour – green to yellow to dark brown at maturity 10-15 stalked microsporophylls with a broad base spirally arranged on the cone axis Microsporophylls are surrounded by ovate scales Each microsporophylls bears 8 or 9 microsporangia in two rows on its abaxial side Before polen dispersal the cone axis elongate to expose the individual sporangia Sporophylls bears stomata on the lower surface and resin ducts in the distal portion Female cones •Develop as inconspicuous axillary bud on shoots of the same year •Round when young, ovoid when mature •Each cone comprises spirally arranged overlapping bracts •Lower and upper part bracts are sterile (do not bear ovules) •The ovules initiate earlier than the initiation of the ovuliferous scales •Later ovuliferous scales overgrows the ovules •Each megasporophyll comprises the bract and ovuliferous scale • In mature cone they fused together at base •Each megasporophyll bears 2-3 ovules on its adaxial surface •After pollination the tip of the ovuliferous grows to meet the scale above thus closing the cavity where the ovule lies •In a cone fertile sporophyll are ten •Seeds are dark brown, compressed, three angled and three winged immature female cones-these are fleshy and green while they develop.. and they will eventually darken to a brown color, and then dry out at maturity. Male cell Part of ovule showing pollen tube Pollen tube near the archegonial complex L.S thru archegonia complex Egg nucleus Stage of fertilization – fusion of male and female gametes A: L.S. of seed showing mature embryo B-D: Progressive stage in the development of seed coat Cupressus Common Name : Mediterranean cypress, common cypress, Italian cypress Family : Cupressaceae Subfamily: Cupressoideae The common name comes from Old French cipres and that from Latin cyparissus which is the latinisation of the Greek Kuparissos Family Cupressaceae •Much branched evergreen trees to shrubs •Plant parts are arranged in opposite and deccusate manner or in whorls of 3 or 4 •Juvenile leaves – linear or spreading (present in seedling and on mature trees) •Leaves are adnate or adpressed hiding the stem •They are glandular or grooved on the abaxial side •Monoecious •Fusion between bract scale and ovuliferous scale prominent •Ovules are erect, 3-20 •Female cones woody at maturity •Seeds may be winged or wingless •Male cone are catkin like, solitary, axillary or terminal •Pollen sacs 3-6 •Pollen lack prothallial cells, wings and sacci 22 genera 150 species 11 are monotypic Half of the genera have distribution in southern hemisphere and half in northern hemisphere Cupressus Distribution cypresses are native to scattered localities in mainly warm temperate regions in the Northern Hemisphere, including North western America, Central America, northwest Africa, the Middle East, the Himalayas, southern China and northern Vietnam 12 speceis of Cupressus distributed C. torulosa is native occurring in the Himalayas from Chamba In H.P to Nepal at an altitude of 1800-2800m Other common planted species in India are C. funebris, C. cashmeriana C. sempervirens, C. macrocarpa and C. lusitanica C. funebris found growing near cemeteries and monasteries and is known as the “chinese weeping cypress” or “funeral cypress” Ecology This species is also tolerant to drought, air currents, wind dust, sleet and atmospheric gases. Its root system is well developed. It succeeds on acid and alkaline soils Morphology •Evergreen tree •15-20m height with pyramidal appearance •Bark thin, fibrous, greyish brown and peels off in long strips •Branching two types horizontal or erect main branches spirally arranged on the main stem drooping side branches – weeping branches •Main branches devoid of leaves; drooping branches bear small, opposite, decussate leaves adpressed to the axis •Lateral pair of leaves folded face to face and facial pair are flattened and grooved in the middle •Two types of leaves Apical zone- small, closely placed, apex acute, serrated margin Mature leaves –longer, brownish and distantly placed •Leaves have prominent midrib and scattered stomata on both the surfaces T.S. of wood Monoecious Reproduction •Male cone 6-7 mm long •Terminally on pendulous branches •Yellow to brown when mature Twig bearing male and female cone Male cone Abaxial Microsporophyll Adaxial •6-8 pairs of microsporophylls •Each bearing 2-6 sporangia abaxially •Sporophylls are arranged opposite, decussate manner on the cone axis •No prothalli cells are formed •Pollen are non winged Develop,ment of spore tetrad and pollen Male cone releasing pollen Female cone •Female cones are 10-12mm long •Axillary on the pendulous branch •Each consists of four pairs of seed scale complexes arranged in an opposite , decussate manner •Lowermost pair is generally sterile •Fertile scales are peltate bears 3-7 ovules or seeds per scale •Fertile seed scale complexes are thick, 4-6 sided with a central pointed structure called boss •Mature cone is woody and peristent •Seeds are dark brown, orbicular, compressed and narrowly winged •The wings of successive seeds overlap