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Gas exchange Photosynthesis Transpiration Xylem – water & mineral transport Water & mineral absorption Phloem – sugar transport Gas exchange – cellular respiration 3 MAIN CELL COMPARTMENTS TISSUE COMPARTMENTS •Symplast – travel via cytosol continum •Apoplast- travel via cell walls & extracellular spaces ABSORPTION OF WATER & MINERALS BY ROOTS Mycorrhizae – •Symbiotic relationship •Surface area for absorption TRANSPORT OF WATER • Root pressure • Guttation • Transpirational pull • Cohesion & adhesion TRANSPIRATIONAL PULL ASCENT OF WATER •SOLAR POWERED •HYDROGEN BONDING •CHARACTERISTICS OF WATER CONTROL OF TRANSPIRATION Photosynthesis – transpiration compromise Stoma open Guard cells Stoma closed Guard cells: inner walls thicker, cellulose microfibrils Guard cells: regulated by uptake and loss of K+ Adaptations to reduce transpiration: thick cuticle, recessed stomata Stoma TRANSLOCATION OF PHLOEM SAP From source (sugar production) to sink (consumes or stores sugar), pressure flow hypothesis Chemiosmotic mechanism for active transport of sucrose PRESSURE FLOW Loading of sugar reduces water potential Absorption of water generates pressure and forces flow Pressure gradient reinforced by the unloading of sugar at the sink Xylem recycles water from sink to source Tapping phloem-sap with the help of an aphid THE AVAILABILITY OF SOIL WATER & MINERALS • Roots hairs increase surface area • Minerals actively transported in, water follows by osmosis Soil Bacteria: Nitrogen fixing & Ammonifying (decomposers) Development of a soybean root nodule Pericycle layer gives rise to secondary roots Root nodules on legumes PARASITIC PLANT CARNIVOROUS PLANTS ANGIOSPERM LIFE CYCLE sporophyte/gametophyte; diploid/haploid FLOWER ANATOMY Complete-all organs Incomplete-lacking 1 or more organs Bisexual – both stamens & carpels Unisexual-one or the other Monoeciouscarpellate & staminate flowers Dioecious-separate plants Angiosperm Gametophytes Pollen grains male female Reduce self - fertilization Genetic Basis of SelfIncompatibility Growth of pollen tube and double fertilization * * Development of a Dicot Embryo Above cotyledons Below cotyledons Embryonic root Development 0f a pea fruit Unique to monocots GERMINATION • • • • Imbibition Release of Gibberellic acid Aleurone enzymes (α amylase) Hydrolysis of endosperm Seed Germination - Radicle emerges 1st a) Cotyledons pulled from soil b) Hypocotyl emerges, cotyledons remain in ground c) Shoot grows up through coleoptile ASEXUAL REPRODUCTION (vegetative reproduction) Fragmentation – separation of a parent plant into parts that reform whole plants Root system of a single parent gives rise to many adventitious shoots Vegetative Propagation - cuttings PLANT RESPONSES TO EXTERNAL SIGNALS Light induced greening of dark sprouted potatoes Grass seedling growing toward light Signal Transduction Pathway (review) TROPISMS • Phototropism – response to light – Stems (positive); Roots (negative) • Gravitropism – response to gravity – Stems (negative); Roots (positive) • Thigmotropism – response to contact – Curling around objects (vines) PHOTOTROPISM CONCLUSION -CHEMICAL SIGNAL PRESENT IN COLEOPTILE TIP STIMULATES GROWTH AS IT PASSED DOWN THE COLEOPTILE -HIGHER CONCENTRATION OF CHEMICAL ON DARKER SIDE CAUSED THE PLANT GROWTH TO CURVE TOWARD LIGHT - NAMED THE CHEMICAL “AUXIN” PLANT HORMONES • Auxins – stem elongation in apical meristems – Fruit maturation, prevents abscission • Cytokinins – cell division in roots, embryos, fruits • Gibberellins – stem elongation in mature regions, fruit development • Abscisic acid – dormancy, stress, abscission • Ethylene – fruit ripening Apical Dominance: •Terminal shoot inhibits lateral buds •Auxin responsible Gibberellins: •Stimulate growth (elongation & division) •Tall spindly plants •Larger seedless grapes Abscisic Acid (ABA) •Seed dormancy -Inhibits germination •Stress -Drought -Winter Leaf Abscisision •Parenchyma cells w/ very thin walls •Change in balance of auxin & ethylene •Aging leaf produces less & less auxin Phytochrome regulation of lettuce seeds Pr ↔ Pfr acts as a switching mechanism that controls various light-induced events Functions as the photoreceptor Links light reception to cellular responses bluish blue-greenish Switched on by Photoperiodic Control of Flowering Short day plants flower when night exceeds the critical dark period Long day plants flower when night is shorter than the critical dark period Root Gravitropism Smaller plant touched 2x/day Rapid turgor movements Response to flooding & oxygen deprivation