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Communities Limiting Factors • Any biotic or abiotic factor that restricts the existence, numbers, reproduction, or distribution of organisms. • Factors that limit one population directly may also have an indirect affect on other populations. Tolerance • The ability of an organism to withstand fluctuations in biotic or abiotic environmental factors. Succession • The orderly, natural changes and species replacements that take place in the communities of an ecosystem. • Occurs in stages. – Each stage has conditions suitable for some organisms but not for others. • Takes decades to centuries to occur. Primary Succession • The colonization of barren land by communities of organisms. • The first species to colonize in barren areas are called pioneer species. • Pioneer species eventually die out. • After some time, the community reaches equilibrium. Climax Community • A stable, mature community that undergoes little or no change. • May last for hundreds of years. Secondary Succession • The sequence of changes that takes place after an existing community is severely disrupted in some way. • Because soil already exists, secondary succession may take less time than primary succession to reach a climax community.