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Why is how species interact with each other important? 1. Interactions can affect the distribution and abundance of species. 2. Interactions can influence evolution. Competition – when two species share a requirement for a limited resource reduces ecological fitness of one or both species Types of ecological competition 1. Intraspecific competition – between individuals of the SAME species contributes to K (carrying capacity) 2. Interspecific competition – between individuals of DIFFERENT species How does interspecific competition affect N? 1 k rats excluded .8 .6 .4 .2 0 # deer mice captured control 1978-80 1988-90 (Heske, E. J., J. H. Brown, and S. Mistry 1994) What is the effect of kangaroo rat competition on deer mice? Competitive exclusion principle If two species have the same niche, the stronger competitor will eliminate the other competitor. “Complete competitors cannot coexist.” Some questions to discuss… Why do kangaroo rats exclude deer mice? Complete competitors cannot coexist What if you put more seeds in the environment? Competitive exclusion is reached more slowly with more resources, but will still happen What if you added another resource to the environment? Stable coexistence requires niche differentiation, these two species are too similar and would still complete directly with one another How does interspecific competition affect N? Lotka-Volterra equation Why isn’t this a complete picture of what happens in nature? A classic interspecific competition experiment two species of Paramecium predict the outcome of interspecific competition P. aurelia P. caudata Gause (1934) What is the niche? set of conditions within which an organism can maintain a viable light intensity population multi-dimensional salinity with as many dimensions as their are limiting conditions okay ecological niche temperature Fundamental niche depends on physical (abiotic) conditions. Realized niche depends on biotic as well as abiotic conditions. What is the realized niche of each barnacle? What is the fundamental niche of each? How can we determine the realized niche of each barnacle? Where do they grow when allowed to compete? growth rate Balanus realized niche low Balanus and Chthamalus Chthamalus realized niche middle high Location in intertidal zone How can we determine the fundamental niche of each barnacle? Removal experiments – remove each species and see where the other grows Balanus alone Balanus fundamental niche growth rate Chthamalus alone Chthamalus fundamental niche low middle high Location in intertidal zone The niche of a species may contract in the presence of a competitor species. This phenomenon leads to resource (niche) partitioning and coexistence among functionally similar species. Streams with only Planaria species A Streams with only Planaria species B Streams with both Planaria species What are the fundamental and realized niches for each species? So…competition is one way in which species can interact with one another. It can be intraspecific (between members of the same species) or It can be interspecific (between members of different species) When invasive species are introduced to an ecosystem, which type of competition do you suspect may result in the reduction of native species populations? Your assignment 1st choose one invasive species that completes directly with a native species (interspecific competition). This should not be a predator/prey interaction. 2nd create an educational “talk” aimed at informing the public about how this invasive species can cause a decrease in (or in some cases disappearance of) native populations. This can be a poster presentation. Include: -A brief biography on each species -An explanation of what ecological competition is -A description of each species fundamental niche -An explanation describing the competitive exclusion -One graph showing how the invasive has impacted native populations