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Ecological Foundations Earth Partnership for Schools Southeast Michigan Institute Suzan Campbell Oakland County n 2009 overview . . . Overview: •Ecology 101 •Landforms • Soils • Regional Landscape Ecosystems • Natural Communities • Internet Resources Activity • Ecosystem Management ecology 101 . . . Ecosystems Ecological Society of America (Christenson et al., 1996) A spatially explicit unit of the Earth that includes all the organisms, along with all the components of the abiotic environment within its boundaries. ecology 101 . . . More than biotic/abiotic components Ecosystems include: •processes •functions •interactions ecology 101 . . . Energy cycle all* energy for life comes from the sun plants capture this energy animals utilize this captured energy from plants energy flow is one way (heat lost) Diagram here ecology 101 . . . Food webs ecology 101 . . . Water cycle earth’s supply of water is fixed stored in aquifers, glaciers, plants land cover affects: infiltration runoff rates erosion flood risk ecology 101 . . . Nitrogen cycle plants can’t use nitrogen in its most abundant form (N2) bacteria and lightning convert it to a useable form (NH4) human activities affect nitrogen’s form, impact in the environment nitrogen is only one of many nutrient cycles ecology 101 . . . Don’t forget the invisible !!! soils provide most Of nutrients needed for plant growth major player in water, nutrient cycles stores water eroded rock, nutrients, decaying organic matter, water, air, billions of living organisms ecology 101 . . . Carrying capacity: maximum pop. size of a particular species that a given habitat can support over a given time period ecology 101 . . . This applies to us, too!!! ecology 101 . . . Whole ecosystems have limits too: need some number of producers, consumers, and decomposers to function must maintain important natural processes and interdependencies no one knows all the limits no one knows which species count We can try to mimic ecosystems that work . . . . . . both native and cultural ecology 101 . . . Succession The orderly and predictable replacement of plants and animals over time The changes are different in different physical settings and under different disturbance regimes. ecology 101 . . . Tolerance “The engine of forest succession is fueled by the relative tolerance of trees to competitive conditions.” Competitive variables: •light •moisture •nutrients •oxygen •space •disturbance Tolerance to low light levels is critical on mesic sites! ecology 101 . . . Shade tolerance Intolerant Tolerant Paper birch Black cherry Yellow birch Basswood Flowering dogwood Aspens Hophornbeam Tuliptree Silver maple Red maple Cottonwood Sycamore White oak Black spruce Beech Pin cherry Black oak Red oak White spruce Sugar maple Tamarack Red ash Hickories Jack Pine Sassafras White ash Red pine Elms Red cedar White pine ecology 101 . . . Gap-phase dynamics In communities of shade-tolerant species, young trees can thrive in the understory until a gap occurs in the canopy •Sugar maple produces prolific seedlings •Beech bends to capture light Sugar maple and beech may not be as competitive in all situations (frequent disturbance, low nutrient, oxygen, etc.) ecology 101 . . . Natural disturbances: ecology 101 . . . Disturbance-adapted communities Fire: Grasslands, barrens, oak-hickory forest, pine forests Flooding: Floodplain forest, swamps, wet meadows, prairies (human) Managed landscapes: Agricultural fields, timber stands, sprawl & urban decay. . . your landscape context . . . Think about your local natural landscape:: • what does it look like? • what are its boundaries? • what plants grow there? • what animals live there? • why has this combination of plants, animals, soils, waterways and landforms ended up together in this space? landforms . . . Michigan’s landscape was shaped by glaciers landforms . . . 14,500 years ago . . . . . . glaciers covered most of the state Interlobate landforms . . . Enormous volumes of meltwater sorted sand and gravel Outwash landforms . . . Where the ice was stagnant and melting in large chunks, kames and eskers were deposited Kame Esker landforms . . . Sometime ice chunks broke off and were buried in debris - when they melted they formed kettles, a kind of depression Kettles landforms . . . Where they paused, moraines were deposited . . . as they melted, lakes formed landforms . . . End moraines landforms . . . End moraines landforms . . . Lake deposits landforms . . . Lake deposits landforms . . . What they left behind . . . Moraines long ranges of hills that trace the original glacial lobes Outwash plains & ice contact features flat plains, kettles, kames & eskers Lakeplains low flat lands with beach ridges soils . . . soils . . . soils . . . Drift: material that has been moved by a glacier Till: unsorted sediments deposited directly by glaciers Stratified drift: sediments that have been sorted by glacial meltwater (outwash, ice contact features) soils . . . Moraines: unsorted (till) – Ranges of hill, soils with mixed particle sizes, often with good water retention, drier if materials are coarser, nutrient-rich, include silts and clays – Support hardwood forests Outwash: sorted sands and gravels – Flat or undulating lands with coarse texture soils, nutrient poor, droughty, fire-prone, can be poorly drained depending on how thick they are and what lies below. – Support grasslands, savannas, oak and pine forests Ice contact features: sorted sands and gravels – Conical hills (kames) or long, linear hills (eskers) – Dry oak and/or pine forest, hillside prairie soils . . . Kettles: ice block depressions – Silts and clays with poor drainage – Lakes, bogs, marshes, swamps Lakeplain: bottom of meltwater lakes – Silty clays and clays – Sandy beach ridges overlying clay – Support hardwood swamps, wet prairies, coastal wetlands on clay, forests, savanna and drier prairie on ridges Bedrock: vary by type of rock – Harsh conditions – Sparse vegetation landscape ecosystems . . . Regional Landscape Ecosystems of Michigan, Minnesota, and Wisconsin: A Working Map and Classification (Dennis A. Albert) . . . the landscape is conceived here as a series of ecosystems, large and small, nested within one another in a hierarchy of spatial sizes. Available online Interactive map interface http://www.npwrc.usgs.gov/resource/habitat/rlandscp.htm landscape ecosystems . . . Sections: based on long-term climate records, * physiography Section IX. Northern Continental M, W & M • continental influenced climate • extremely cold in winter • lake effect precipitation along Lake Superior Section VIII. Northern Lake Influenced Upper M & W • lake moderated temperatures • lake effect precipitation along Lake Superior Section VII. Northern Lake Influenced Lower Michigan • lake moderated temperatures • lake effect snow near shorelines • interior has greatest weather extremes Section VI. Southern Lower Michigan • longest growing season • lake moderated temperatures • more warm humid air/less cold dry air * physiography – form of the land and parent material landscape ecosystems . . . Sections: based on long-term climate records, * physiography Section IX. Northern Continental M, W & M • continental influenced climate • extremely cold in winter • lake effect precipitation along Lake Superior Section VIII. Northern Lake Influenced Upper M & W • lake moderated temperatures • lake effect precipitation along Lake Superior Section VII. Northern Lake Influenced Lower Michigan • lake moderated temperatures • lake effect snow near shorelines • interior has greatest weather extremes Section VI. Southern Lower Michigan • longest growing season • lake moderated temperatures • more warm humid air/less cold dry air * physiography – form of the land and parent material landscape ecosystems . . . Sub-sections & sub-subsections: * based on long-term climate records, physiography Section VI. Southern Lower Michigan Sub-section VI.1. Washtenaw landscape ecosystems . . . Sub-subsections: based on physiography (land form/parent material) - because it controls fluxes of radiation and moisture and thereby strongly determines the pattern of soil, microclimate, and vegetation. Section VI. Southern Lower Michigan Sub-section VI.1. Washtenaw • Sub-sub-section VI.1.1 Maumee Lakeplain • Sub-sub-section VI.1.2 Ann Arbor Moraines • Sub-sub-section VI.1.3 Jackson Interlobate thinking locally . . . Natural communities Background so far: •natural process disturbances •landform •soil •climate We’ll be adding •biota thinking locally . . . Natural communities recurrent interacting assemblage of climate, landform, soil, native plants, animals, and dynamic processes at a local scale thinking locally . . . Natural communities recurrent interacting assemblage of climate, landform, soil, native plants, animals, and dynamic processes at a local scale identified by dominant vegetation natural communities . . . In a highly altered landscape: How do we know which assemblages of •climate, •landform, •soil, •plants & animals •dynamic processes are natural ? natural communities . . . Community types forest grassland and savanna open wetlands “primary” communities natural communities . . . Presettlement vegetation map (Comer et al., 1995) based on surveyors records from the 1800s supplemented by years of field work by MNFI staff, historic literature and museum records a “best guess” – not infallible, but still very useful circa 1800 vegetation . . . circa 1800 vegetation . . . Northern coniferous forest Eastern deciduous forest circa 1800 vegetation . . . northern hardwoods, boreal forest, pine forests, conifer swamps northern hardwoods, peatlands, alvar, cedar swamps cedar swamps northern hardwoods pine barrens pine forests oak - pine barrens beech maple forests oak hickory forests savannas, prairies natural communities . . . Michigan’s natural communities: abstracts . . . Abstracts: natural communities, plants and animals element occurrences . . . MNFI’s element occurrence lists: By county or watershed: forest . . . Mesic southern forest Beech Sugar maple forest . . . Mesic southern forest occurs on moraines, old beach ridges* rich, moist, well-drained soils shade tolerant species (reproduce in shade) abundant spring flora vernal pools (29 amphibian spp., 8 reptile spp.) “gap phase dynamics” in regeneration small-scale wind storms, ice storms, primary disturbance * south of the tension zone forest . . . Mesic southern forest Critical processes: Gap phase dynamics • small canopy gaps create temporary increase in light, nutrients and water • allow regeneration of shade tolerant maples forest . . . Dry-mesic southern forest White oak Beech Sugar Blackmaple oak forest . . . Dry, dry-mesic southern forest occurs principally on glacial outwash, coarsetextured moraines, sandy lakeplain & dunes sandy loam and loam soils are slightly acid to neutral shade intolerant species – fire allows regeneration of shade intolerant oak/ reduces shade tolerant invaders forest . . . Dry, dry-mesic southern forest Critical processes: Fire-dependent system • historically, oak openings, barrens, prairie – shifting matrix • maintained by frequent ground fires, infrequent crown fires • suite of species, related communities that benefit from fire oak forests oak savanna dry sand prairie forest . . . Floodplain forest Cottonwood Sycamore Silver maple forest . . . Floodplain forest occurs next to large rivers, frequently in sandy glacial outwash, sand lakeplain* fertile, seasonally saturated soils – mineral at water’s edge, may be organic in back swamp. shade intolerant species along water’s edge complex zonation forest . . . Floodplain forest Critical processes: •Flooding and windthrow frequent • sunlight penetrates along water •Bank-cutting on outer bank, deposition on inner edge forest . . . Wet-mesic flatwoods One of the most distinctive communities in southern Michigan is found on the clayey and seasonally wet lake plain of Belle Isle, located in the Detroit River, Wayne County. Beech Forests there include a unique community of the rare species shumard oak, pumpkin ash and shellbark hickory, together with silver maple, red ash, pin and swamp white oaks, and hawthorns. BV Barnes, Michigan Trees, 2004 Sugar maple forest . . . Wet-mesic flatwoods still being characterized – little known/little left occurs on clay lakeplain or shallow sand over clay heavy, poorly drained soils, seasonally high water table moderately shade tolerant or shade intolerant species – lots of oaks and ashes, silver maple a number of more southern species found here - several rarities still present forest . . . Wet-mesic flatwoods Critical processes: wind throw, seasonally high water table • larger gaps permit persistence of shade intolerant species • seasonally high water tables keep out sugar maple, beech grassland & savanna . . . Lakeplain oak opening Needlegrass Little bluestem open wetlands . . . Emergent marsh open wetlands . . . Emergent marsh occurs in shallow waters at the edge of inland lakes and streams soils are saturated organic mucks shade intolerant species requires periodic flooding to exclude invasion by woody plants open wetlands . . . Emergent marsh Critical processes: Flooding and drawdown • flooding excludes woody invaders • drawdown exposes seedbank, so that light-sensitive annual seeds can germinate • wetland seed remains viable for over 60 years open wetlands . . . Bog open wetlands . . . Bog occurs in depressions in outwash, kettles in end moraine or pitted outwash – may form mat around perimeter of open water soils are extremely acidic peat, may have minerotrophic variants south of the transition zone low nutrient availability can occur in complexes with prairie fen, relict conifer swamp and poor conifer swamp open wetlands . . . Bog Critical processes: Rainwater-fed • minimal input from groundwater • sphagnum acidifies water primary community . . . Limestone pavement lakeshore primary community . . . Limestone pavement lakeshore occurs on bedrock soils are undeveloped except in cracks extremely harsh growing conditions, sparse vegetation primary community . . . Open dunes primary community . . . Open dunes deposited by wind soils are pure sand (parent material) shade intolerant species – adapted to constantly shifting substrate (common milkweed belongs here) Internet resources . . . We’ll be using: Regional landscape ecosystems doc Quaternary geology map Circa 1800 map Element occurrence data Community abstracts planning . . . How do we ensure the conservation of a living organism? protect the lands they need to survive manage ECOSYSTEMS (interacting organisms and their environment) planning . . . Conserve all the pieces and processes – representation (some of every ecosystem) planning . . . Conserve all the pieces and processes – representation (some of every ecosystem) – redundancy (how much is enough?) planning . . . Conserve all the pieces and processes – representation (some of every ecosystem) – redundancy (how much is enough?) – resilience (ability to adapt to changing conditions and stresses) planning . . . Conserve all the pieces and processes – representation (some of every ecosystem) – redundancy (how much is enough?) – resilience (ability to adapt to changing conditions and stresses) Consider – size/shape/configuration on landscape planning . . . Conserve all the pieces and processes – representation (some of every ecosystem) – redundancy (how much is enough?) – resilience (ability to adapt to changing conditions and stresses) Consider – size/shape/configuration on landscape planning . . . Conserve all the pieces and processes – representation (some of every ecosystem) – redundancy (how much is enough?) – resilience (ability to adapt to changing conditions and stresses) Consider – size/shape/configuration on landscape – connectivity (corridors for dispersal, feeding, etc.) planning . . . Conserve all the pieces and processes – representation (some of every ecosystem) – redundancy (how much is enough?) – resilience (ability to adapt to changing conditions and stresses) Consider – size/shape/configuration on landscape – connectivity (corridors for dispersal, feeding, etc.) planning . . . Conserve all the pieces and processes – representation (some of every ecosystem) – redundancy (how much is enough?) – resilience (ability to adapt to changing conditions and stresses) Consider – size/shape/configuration on landscape – connectivity (corridors for dispersal, feeding, etc.) – context (threats, adjacent land uses) planning . . . Conserve all the pieces and processes – representation (some of every ecosystem) – redundancy (how much is enough?) – resilience (ability to adapt to changing conditions and stresses) Consider – size/shape/configuration on landscape – connectivity (corridors for dispersal, feeding, etc.) – context (threats, adjacent land uses) – condition (viable populations, functions intact) planning . . . Coarse Filter/Fine Filter approach Coarse filter (core areas): planning . . . Coarse Filter/Fine Filter approach Coarse filter (core areas): – representative examples of all native and valued community types Prairie fen planning . . . Coarse Filter/Fine Filter approach Coarse filter (core areas): – representative examples of all native and valued community types – allow or mimic natural disturbances Flooding Prairie fen Fire planning . . . Coarse Filter/Fine Filter approach Coarse filter (core areas): – representative examples of all native and valued community types – allow or mimic natural disturbances – captures many elements of biodiversity Flooding Prairie fen Fire planning . . . Coarse Filter/Fine Filter approach Coarse filter (core areas): – representative examples of all native and valued community types – allow or mimic natural disturbances – captures many elements of biodiversity Fine filter (smaller patches): – viable populations of vulnerable species Yellow rail planning . . . Coarse Filter/Fine Filter approach Coarse filter (core areas): – representative examples of all native and valued community types – allow or mimic natural disturbances – captures many elements of biodiversity Fine filter (smaller patches): – viable populations of vulnerable species – capture things that fall through the cracks Yellow rail Lake sedge planning . . . Coarse Filter/Fine Filter approach Coarse filter (core areas): – representative examples of all native and valued community types – allow or mimic natural disturbances – captures many elements of biodiversity Fine filter (smaller patches): – viable populations of vulnerable species – capture things that fall through the cracks Corridors – link core areas and patches planning . . . Coarse Filter/Fine Filter approach Coarse filter (core areas): – representative examples of all native and valued community types – allow or mimic natural disturbances – captures many elements of biodiversity Fine filter (smaller patches): – viable populations of vulnerable species – capture things that fall through the cracks Corridors – link core areas and patches – allows dispersal, migration, large ranging organisms planning . . . Coarse Filter/Fine Filter approach Coarse filter (core areas): – representative examples of all native and valued community types – allow or mimic natural disturbances – captures many elements of biodiversity Fine filter (smaller patches): – viable populations of vulnerable species – capture things that fall through the cracks Corridors – link core areas and patches – allows dispersal, migration, large ranging organisms Thoughtful management of the lands in between – maximize biodiversity conservation, where possible The Principles: • • • • • • • Goals – what are you managing for? Sound ecological models and understanding Complexity and connectedness Dynamic character of ecosystems Context and scale Humans as ecosystem components Sustainability (carrying capacity) • Use best available information and implement strategy Learning is • Adaptability and accountability fundamental to the process! – did it work? know your land . . . Questions? ecosystem management . . . What should we be doing? Congratulations!! You just got a paying job to manage your favorite natural area, wild area, recreational area, green way, park, open space… ecosystem management . . . Ecosystem Management: • • • • DEFINE YOUR GOALS!!!! Ecosystem integrity and function Long term sustainability (carrying capacity) Biodiversity conservation (pieces and connections) • Sustaining the system = sustains those things we desire • Integrate social and economic factors ecosystem management . . . Sustainability Economic Social Ecological ecosystem management . . . economic ecological social Understanding ecosystems helps us set appropriate management objectives. composition (pieces) structure (organization) function (how it works) Ecosystems have limits!! ecosystem management . . . Sustainability Economic Redefine goals if necessary. Social Ecological Speak up about ecosystem contstraints. processes, functions, interactions . . . Not just biotic/abiotic components: • interdependencies that allow things we value to thrive Eastern prairie fringed orchid • mycorrhizae (root-fungi) • flooding/fire dependent • emit odor at night Pandorus sphinx (Eumorpha pansorus) Photo by Dave Cuthrell, MNFI Where does the moth live? The sum is greater than its parts! Ecosystem Management... The Principles: • Goals – what are you managing for? • • • • • • Sound science and understanding Complexity and connectedness Dynamic character of ecosystems Context and scale Humans as ecosystem components Sustainability (carrying capacity) • Best available information and implement strategy • Adaptability: did it work? Learning is fundamental! forest . . . Dry, dry-mesic southern forest White oak Black oak forest . . . Dry, dry-mesic southern forest occurs principally on glacial outwash, coarse-textured moraines, sandy lakeplain & dunes sandy loam and loam soils are slightly acid to neutral shade intolerant species – fire allows regeneration of shade intolerant oak/ reduces shade tolerant invaders forest . . . Dry, dry-mesic southern forest Critical processes: Fire-dependent system • historically, oak openings, barrens, prairie – shifting matrix • maintained by frequent ground fires, infrequent crown fires • suite of species, related communities that benefit from fire oak forests oak savanna dry sand prairie Forrest B.H. Brown, 1917: Probably the most obvious characteristic of the forest vegetation as a whole is the extreme variety of species composing many of the associations and the general dissimilarity between it and the usual upland type lying westward and mostly outside of the county. One may clearly observe this transition in going from Ann Arbor to Detroit. The change is abrupt and takes place a short distance west of Ypsilanti or some twenty miles west of the Detroit River, where the rolling morainal topography changes to the level glacial lake basin of which Wayne County, except the small northwest portion is a part. ranking elements . . . Global & State Ranks G1 – globally critically imperiled G2 – globally imperiled G3 – vulnerable G4 – apparently secure (uncommon, not rare) G5 – demonstrably widespread, abundant and secure S1 – critically imperiled within the state S2 – imperiled within the state S3 – vulnerable S4 – apparently secure (uncommon, not rare) S5 – widespread, abundant and secure within the state