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Chapter 7 Community Interactions Let’s get ready to ruuuuumble! Community • Community: populations of all species living and interacting in an area at a particular time Four Characteristics of Community Structure • Physical Appearance: size and distribution of its population and species • Species Diversity/Richness: number of different species • Species Abundance: number of individuals of each species • Niche Structure: number of niches, how they compare, and how they interact Differing Physical Appearances • Patch Effects: most large communities usually consist of a mosaic of vegetation “patches” • Edge Effects: differences in physical appearance at boundaries between ecosystems Species Diversity • Species Rich Environments – – – – – Tropical Rain Forests Coral Reefs Deep Sea Large Tropical Lakes Tend to have high species diversity but low species abundance • Factors that Affect Diversity – Latitude (terrestrial) – distance from equator – Depth (aquatic) – Pollution (aquatic) Species Abundance • Determined by: – Rate at which new species immigrate – Rate at which species become extinct Ecological Niche vs. Habitat Niche: role an organism plays in an ecosystem •niche is like an "occupation“ – a species’ interactions with habitat and other organisms (their role in food web) Habitat: actual location where an organism lives •habitat is like an "address" 7-2 GENERAL TYPES OF SPECIES GENERAL TYPES OF SPECIES •Generalist Species •Specialist Species •Native Species •Nonnative Species •Indicator Species •Keystone Species Generalists vs. Specialists •Generalist Species have broad niches, can live many places, use a variety of resources e.g., dandelions, cockroaches, coyotes, humans •Specialist Species have narrow niches, live only in specific places e.g., spotted owls, giant pandas © Brooks/Cole Publishing Company / ITP Native vs. Nonnative Species • Native Species: species that normally live & thrive in a particular ecosystem • Nonnative Species: •also called exotic, invasive, or alien species •originate in other ecosystem •deliberate or accidental introduction by humans causes problems •may thrive and crowd out native species Emerald Beetle Larva under bark of ash tree Why Purple??? The Case of the Killer Bees • 1957 Brazil imported wild African bees to help increase honey production • Displaced domestic honeybees • Actually reduced honey production • Moved north in Central America • Established populations in Texas, Arizona, New Mexico, Puerto Rico, and California Stink Bugs! Should we release Asian Wasps to neutralize the stink bug population??? Snakehead Fish CANE TOADS! The Simpsons! • Bart vs. Australia Indicator Species • Indicator Species: species that serve as early warnings that a community or ecosystem is being damaged •Birds are good indicator species •Found everywhere •Respond to environmental change quickly •northern spotted owls are indicators of healthy old– growth forest •Fish are good indicator species in aquatic ecosystems (i.e. trout) “The loss of a keystone species is like a drill accidentally striking a power line. It causes lights to go out all over.” Keystone Species – E.O. Wilson Keystone Species: species that play a critical role in an ecosystem •Ex: sea otters are keystone species because they prevent sea urchins from depleting kelp beds •Ex: flying foxes are keystone species because they pollinate & disperse tropical trees such as durian 7-3 SPECIES INTERACTIONS: COMPETITION AND PREDATION Species Interactions • The effects of one species on another may be negative, positive, or neutral • Five kinds of interactions: •Interspecific competition •Predation •Parasitism •Mutualism •Commensalism Intra- vs. Inter- specific competition • Intraspecific Competition – members of the SAME species compete for resources •Interspecific Competition – members from 2 or more DIFFERENT species compete for resources •When two or more species use the same limited resource (food, space, etc.) they may adversely affect each other •niche overlap •Ex: fire ants & native ants in North America No Competition High Relative population density Paramecium aurelia Low Paramecium caudatum 0 2 4 6 8 10 12 Days Each species grown alone 14 16 18 Interspecific Competition High Relative population density Paramecium aurelia Paramecium caudatum Low 0 2 4 6 8 10 12 Days Both species grown together 14 16 18 Resource Partitioning Species with similar resource requirements can coexist because they use limited resources: •at different times •in different ways •in different places © Brooks/Cole Publishing Company / ITP Resource Partitioning Where are the 2 species competing? Why is there no competition here? Resource Partitioning © Brooks/Cole Publishing Company / ITP Resource Partitioning Five species of insect–eating warblers coexist in spruce forests of Maine: •feed in different portions of trees •consume somewhat different insects © Brooks/Cole Publishing Company / ITP Warblers Predation Members of one species (predator) feed on another species (prey); •+ / – •Ex: lion feeding on zebra © Brooks/Cole Publishing Company / ITP Predator-Prey Relationship • What do predators do to increase their chances of getting a meal? • What do prey do to avoid being eaten? Predator-Prey Relationship • Predators get better at catching prey • Prey get better at avoiding capture • Similar to an “arms race” – Ex: During the Cold War, the US and the USSR tried to intimidate the other with bigger and better weapons Predators • Pursuit – Faster (cheetahs) – Better eyesight (eagles) – Hunting in packs (wolves) • Ambush – Camouflage (praying mantis) – Mimicry (alligator snapping turtles) Prey • Camouflage • Ex: walking stick • Warning colors • Ex: poison dart frog • Mimicry • Ex: king snake • Behavior • Ex: blowfish • Chemical warfare • Ex: skunk 7-4 SPECIES INTERACTIONS: PARASITISM, MUTUALISM, AND COMMENSALISM Parasitism One organism (parasite) lives on part of another organism (host) •+ / – •Ex: flea living on a dog •Ex: tongue eating parasite © Brooks/Cole Publishing Company / ITP Mutualism Two species interact in a way that benefits both •+ / + •Ex: lichens (algae & fungi) •Ex: clownfish & anemones •Ex: ants & acacias WITH FRONDS LIKE THESE, WHO NEEDS ANEMONES??? © Brooks/Cole Publishing Company / ITP Commensalism One organism benefits from another, but neither helps nor harms the other organism •+ / 0 •Ex: epiphyte growing on a tree © Brooks/Cole Publishing Company / ITP Species Interactions • the effects of one species on another may be negative, positive, or neutral • five kinds of interactions: POPULATION A COMPETITION PREDATION PARASITISM COMMENSALISM MUTUALISM POPULATION B 7-5 ECOLOGICAL SUCCESSION: COMMUNITIES IN TRANSITION Ecological Succession Succession: gradual & fairly predictable change in species composition over time © Brooks/Cole Publishing Company / ITP Primary Succession © Brooks/Cole Publishing Company / ITP Primary Succession • Type of succession that occurs where there was no ecosystem before • Occurs on rocks, cliffs, and sand dunes • Pioneer species: the first organism to colonize any newly available area and begin the process of ecological succession Primary Succession Primary Succession: gradual establishment of biotic communities in an area where no life existed before •Ex: succession on newly formed islands & after the retreat of a glacier •Early Communities: lichens & mosses colonize bare rock •Mid Communities: small herbs & shrubs colonize •Late Communities: tree species colonize © Brooks/Cole Publishing Company / ITP Lichens Mosses Parking Lot Secondary Succession © Brooks/Cole Publishing Company / ITP Secondary Succession Secondary Succession: gradual reestablishment of biotic communities in an area where a biotic community was previously present •Ex: "old field succession" © Brooks/Cole Publishing Company / ITP Secondary Succession: Mount St. Helens • Erupted in 1980 • 44,460 acres were burned and flattened • After the eruption, plants began to colonize the debris Disturbance Disturbance: a distinct event that disrupts an ecosystem or community •disturbance initiates secondary succession •Natural disturbances: fires, hurricanes, tornadoes, droughts, & floods •Human-caused disturbances: deforestation, overgrazing, plowing Intermediate Disturbance Hypothesis: Moderate disturbances in communities promote greater species diversity than small of major disturbances Climax Community • Climax community: the final and stable community in an ecosystem • Will continue to change in small ways, but will remain primarily the same over time (unless disturbed) The Circle of Life in Secondary Succession 7-6 ECOLOGICAL STABILITY AND SUSTAINABILITY Stability - complex networks of positive and negative feedback loops • maintained only by constant dynamic change in response to changing environmental conditions Ecosystem Stability • Inertia: the ability of an ecosystem to resist being disturbed (also called persistence) • Constancy: ability of a living system (population) to keep its numbers w/in the limits imposed by available resources • Resilience: the ability of an ecosystem to “bounce back” after it has been disturbed Precautionary Principle • We should try to prevent potential harm to an ecosystem even though we don’t understand all of the cause-and-effects • “better safe than sorry”