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Biological Communities • Community: all the organisms that live together in a specific place – ______________________ – _______________________ – _______________________ – ______________________ • Communities can be characterized either by their constituent species or by their properties – Species richness: the number of species present – Primary productivity: the amount of energy produced 1 Biological Communities • Two views of structure and functioning of communities – Individualistic concept: H.A. Gleason; a community is nothing more than an _____________________________that happen to occur together at one place – Holistic concept: F.E. Clements: a community is an integrated unit; _______________________________more than the sum of its parts 2 Biological Communities Most ecologists today favor ______________ _________________ • In communities, species respond independently to changing environmental conditions • Community composition changes gradually across landscapes 3 Biological Communities • Abundance of tree species along a moisture gradient in the Santa Catalina Mountains of Southeastern Arizona • Each line represents the abundance of a different tree species • Community composition ____________________ ____________________ 4 Biological Communities • Sometimes the abundance of species in a community does change geographically in a synchronous pattern • ________________: places where the environment changes abruptly 5 Ecological Niche • Niche: the total of all the ways an organism uses the resources of its environment – _______________________________ – _______________________________ _______________________________ ________________________________ • Fundamental niche: the entire niche that a species is ____________________________________, based on physiological tolerance limits and resource needs • Realized niche: __________________ set of environmental conditions, _____________________or _____________________of other species, in which the species can establish a stable population 6 Billock Ecological Niche • Interspecific competition: occurs __________________ ______________________________ and there is not enough resource to satisfy both 7 Ecological Niche • Other causes of niche restriction – ____________________ absence or presence • Plant species – Absence of pollinators – Presence of herbivores Billock Billock 8 Ecological Niche Principle of competitive exclusion: ___ __________________________________ __________________________________ __________________________________ __________________________________ • G.F. Gause’s classic experiment on competitive exclusion using three Paramecium species shows this principle in action 9 10 Ecological Niche • Niche overlap and coexistence • Competitive exclusion redefined: no two species can occupy the same niche indefinitely _________________________ • Species may divide up the resources, this is called _____________________ • Gause found this occurring with two of his Paramecium species 11 • Resource partitioning is often seen in _________________ _________________ that occupy the same geographic area • Thought to result from the process of _________________ _________________ Resource partitioning among sympatric lizard species 12 • Character Ecological Niche displacement: differences in morphology evident between sympatric species – May play a role in adaptive radiation 13 Ecological Niche • Detection of interspecific competition can be difficult – If resources not limited there may be no competition – Small versus large population size – May be environmental conditions that cause the decline of a species, not competition 14 Ecological Niche Detecting interspecific competition 15