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Post Graduate Diploma in Environment Education Unit 7 Principles pertaining to limiting factors and ecological assessment for sustainability 1.Liebig’s law of the minimum 2.Shelfords’s law of tolerance 3.Regulatory factors 4.Ecological indicators Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 1 Post Graduate Diploma in Environment Education Unit 7 Principles pertaining to limiting factors and ecological assessment for sustainability Liebig’s law of the minimum: Growth of a plant is dependent on the amount of foodstuff which is presented to it in minimum quantity. This law of the minimum was first expressed by Justus Liebig in 1840. This law of the minimum is less applicable under ‘transientstate’ conditions when the amounts, and hence the effects, of many constituents are rapidly changing. Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 2 Post Graduate Diploma in Environment Education Unit 7 Principles pertaining to limiting factors and ecological assessment for sustainability Shelfords’s law of tolerance: The existence, abundance, and distribution of a species in an ecosystem are determined by whether the levels of one or more physical or chemical factors fall within the range tolerated by that species. In other words, there are minimum and maximum limits for physical conditions (such as temperature) and concentrations of chemical substances-called tolerance limits-beyond which no members of a particular species can survive. Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 3 Post Graduate Diploma in Environment Education Unit 7 Principles pertaining to limiting factors and ecological assessment for sustainability Shelfords’s law of tolerance This concept of the limiting effect of maximum as well as minimum was incorporated into the ‘law’ of tolerance by V. E. Shelford in 1913. Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 4 Post Graduate Diploma in Environment Education Unit 7 Principles pertaining to limiting factors and ecological assessment for sustainability Limiting Factors The presence and success of an organism or a group of organisms depends upon a complex of conditions. Any condition which approaches or exceeds the limits of tolerance is said to be a limiting condition or a limiting factor. Fig: A Model summarizing general principles of limiting factors (source Odum, 1971) Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 5 Post Graduate Diploma in Environment Education Unit 7 Principles pertaining to limiting factors and ecological assessment for sustainability Physical Factors of Importance as Limiting Factors TEMPERATURE: 1. Life can exist only within -200deg to 100 deg C. 2. Most species are restricted to a narrower range of temperature 3. Aquatic organisms have narrower range of tolerance than equivalent land animals 4. Organisms which are subjected to temperature variations tend to be depressed, inhibited or slowed down by constant temperature Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 6 Post Graduate Diploma in Environment Education Unit 7 Principles pertaining to limiting factors and ecological assessment for sustainability Physical Factors of Importance as Limiting Factors LIGHT: The quality of light, the intensity, and the duration are known to be important factors of light. Both plants and animals respond to different quality of light. Individual plants as well as communities adapt to different light intensities by becoming ‘shadeadapted’. Or ‘sun adapted’. Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 7 Post Graduate Diploma in Environment Education Unit 7 Principles pertaining to limiting factors and ecological assessment for sustainability Physical Factors of Importance as Limiting Factors WATER: Rainfall distribution over the year is an extremely important limiting factor for organisms. Humidity has a daily rhythm (high at night, low during the day), and has special role in modifying the effects of temperature, hence regulates the activities of organisms and in limiting their distribution. Evapo-transpiration is also an important limiting factor. Desert plants expose only green buds or stems. Dew , in areas of low rainfall, is a vital contributor to precipitation. Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 8 Post Graduate Diploma in Environment Education Unit 7 Principles pertaining to limiting factors and ecological assessment for sustainability Physical Factors of Importance as Limiting Factors ATMOSPHERIC GASES: The concentration of carbon di-oxide an oxygen is limiting to many higher plants. BIOGENIC SALTS (MACRONUTRIENTS & MICRONUTRIENTS: Nitrogen and phosphorous salts are of major importance to ecologists as limiting factors. CURRENTS & PRESSURES Currents in water not only influence the concentration of gases and nutrients, but act directly as limiting factors. Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 9 Characteristics of ecological indicators 1. In general, steno species make much better indicators than eury species. Such species are often not the most abundant ones in the community. 2. Large species usually make better indicators than small species. A large and more stable biomass and standing crop can be supported with a given energy flow. The turnover rate of small organisms may be so great that the particular species present at any time may not be very instructive as an ecological indicator. Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 10 Characteristics of ecological indicators 3. Before relying on as single species or groups of species as indicators, there should be abundant field evidence. 4. Numerical relationships between species, populations, and whole communities often provide more reliable indicators than single species. Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 11 If we love our children, we must love the country with tender care and pass it on, diverse and beautiful, so that our children can hear a Danphe call in the Himal, feel peace in hills and mountains with forest of Rhododendron, can watch a one horned Rhinoceros in Terai and can find joy in being alive. -Subodh Sharma Subodh Sharma Dr.nat.tech.; Aquatic Ecology Centre, P.O. Box: 6250, [email protected]. 12