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Name ____________________________________ Date ____________ Period _____ Agents of Evolutionary Change Raven Ch. 21 Big Idea 1: THE PROCESS OF EVOLUTION DRIVES THE DIVERSITY AND UNITY OF LIFE. It is not the strongest of the species that survives, nor the most intelligent that survives. It is the one that is the most adaptable to change. ― Charles Darwin (1859) Essential Knowledge: Natural selection is a major mechanism of evolution Natural selection acts on phenotypic variations within populations. Evolutionary change is also driven by random processes. Biological evolution is supported by scientific evidence from many disciplines including mathematics Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. Phylogenetic trees and cladograms are graphical representations (models) of evolutionary history that can be tested Speciation and extinction have occurred throughout the Earth’s history. Speciation may occur when two populations become reproductively isolated from each other. Populations of organisms continue to evolve. There are several hypotheses about the origin of life on Earth, each with supporting scientific evidence. Scientific evidence from many different disciplines supports models of the origin of life. Populations Evolve! o Natural Selection__________________________________________________________ ______________________ results in: o differential survival o differential reproductive success o “survival of the fittest” who bears more offspring _______________________ evolve __________________ makeup of population changes over time favorable traits (_____________________________) become more common Individuals ___________________ evolve… Individuals __________________ or ______________________… Individuals __________________________ or ________________… Individuals ______________________ evolve… Individuals are ______________________ ___________________________ evolve 1 o Variation & Natural Selection ________________________ is the raw material for natural selection o there have to be _____________________ within population o some individuals must be __________________ than others ________________________- Survival & Reproductive success o o individuals with one phenotype leave more surviving offspring Where does Variation come from? _______________________ _________________ changes to _________________ errors in mitosis & meiosis (cell division) environmental damage _________________ mixing of alleles ______________________ of alleles o new combinations = new phenotypes spreads variation Examples: new arrangements in every offspring offspring inherit traits from parent _____________________________________________________, _____________________________________________________, _____________________________________________________, _____________________________________________________, How can the alleles within a population change (evolution)? 5 Agents of evolutionary change 1. ___________________________________ 2. ___________________________________ 3. ___________________________________ 4. ___________________________________ 5. ___________________________________ 2 1. Mutation & Variation Mutation creates __________________________________ o Mutation rates in _________________________ are __________________ (1 in 100,000 division) o But are constantly appearing in _________________________ Mutation changes __________________________________________. o As a result, may change: Protein amino acid sequence may change protein _________________________? may change protein _________________________? changes in protein may change __________________ & therefore change __________ 2. Gene Flow Movement of ___________________ & ____________________ in & out of populations seed & pollen distribution by __________________________ __________________ of animals 1. sub-populations may have different allele frequencies 2. causes _____________________________ across regions 3. ________________________ differences between populations 3. Non-random mating ______________________________ 4. Genetic drift Effect of __________________ events o founder effect o small group splinters off & starts a new colony bottleneck some factor (disaster) reduces population to small number & then population recovers & expands again 3 5. Natural selection Differential ___________________ & _________________ due to changing environmental conditions climate change food source availability predators, parasites, diseases toxins combinations of _______________________ that provide “_________________” ______________ in the population _________________________ evolutionary change 4 Measuring Evolution in Populations through Hardy-Weinberg Raven Ch. 21 Populations & gene pools Concepts: a ____________________________ is a localized group of ___________________ individuals gene pool is collection of ______________________ in the population remember difference between alleles & genes! ________________________________ is how common is that allele in the population how many _____ vs. _____ in whole population Evolution of populations Evolution = ________________________________________________________________________ Hypothetical: what conditions would cause allele frequencies to NOT change? ________________________________________________ _____________________________ all agents of evolutionary change 1. very large population size (no _______________________________) 2. no migration (no __________________________ in or out) 3. no _____________________________ (no genetic change) 4. _______________________________ (no sexual selection) 5. no ___________________________________ (everyone is equally fit) Hardy-Weinberg equilibrium _____________________________________________________________ preserves allele frequencies Serves as a model (________________________ hypothesis) natural populations ________________________ in H-W equilibrium useful model to ___________________________ if forces are acting on a population measuring evolutionary change Hardy-Weinberg theorem Counting _____________________________________ assume ____________ alleles = _______, _______ frequency of dominant allele (B) = ________ frequency of recessive allele (b) = ________ frequencies must add to 1 (100%), so: _____ + _____ = 1 5 Counting ____________________________________ ( _____________________________) frequency of ______________________: p x p = _____ frequency of ______________________: q x q = _____ frequency of ______________________: (p x q) + (q x p) = ______ frequencies of all individuals must add to 1 (100%), so: __________________________ H-W formulas: Alleles: H-W Problem: Individuals: Population: 100 cats, 84 black, 16 white How many of each genotype? Must assume population is in H-W equilibrium Sample Data: How do you explain the data? Sample Data: How do you explain the data? 6 Application of H-W principle _______________________________ inherit a mutation in gene coding for _________________________ oxygen-carrying blood protein recessive allele = ___________ normal allele = ___________ low oxygen levels causes RBC to sickle breakdown of RBC clogging small blood vessels damage to organs often __________________ Sickle cell frequency High frequency of ___________________________ 1 in 5 in Central Africans = HbHs unusual for allele with severe detrimental effects in homozygotes 1 in 100 = HsHs usually die before reproductive age Why is the Hs allele maintained at such high levels in African populations? Answer: ________________________________ In tropical Africa, where malaria is common: ________________________________ (normal) die or reduced reproduction from malaria: HbHb ________________________________ die or reduced reproduction from sickle cell anemia: HsHs ________________________________ are relatively free of both: HbHs survive & reproduce more, more common in population Example: 7