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The LEAF Competencies: Identify the parts of the Leaf and its functions and classifications Pre Activity: 1. Get a sample of a leaf (any leaf or leaves with stalk) 2. Recall the parts of the leaf THE LEAF LEAF – expanded lateral extension of the stem arising at the node with a bud on its axil. 2 types: a. microphyll – lycopods with one vascular strand. b. megaphyll – complex venation The Plant Body: Leaves • FUNCTION OF LEAVES – Leaves are the solar energy and CO2 collectors of plants. – In some plants, leaves have become adapted for specialized functions. External Parts: Petiole – stalk which connects the lamina to the stem. Function: act as passage of substances between leaf and stem. leaf without petiole – sessile with petiole – petiolate peltate – where petiole is attached some distance from the leaf. 2. Leaf Blade or Lamina – expanded portion of the leaf that varies on the tips, bases and margins. Function: where photosynthesis takes place. 3 parts: a. tip – apex b. side – margin c. posterior – base 3. Stipules – leafy appendages arising from the base of the petiole. with stipule – stipulate without stipule – exstipulate function: for photosynthesis/ protection 4. Midrib – large vessel serve as an extension of petiole. Function: conduction and support of leaf tissues. 5. Netwrok of Vein and Veinlets – also called Nervules- smaller branches of vessels running on leaf blade. for conduction and transport of materials. Classification of Leaves: 1. Venation – arrangement of veins *parallel – one sided *netted/reticulate – close network *obsolete/wanting – too obscured because they are too thick. • common to dicots and some nonflowering plants. – Pinnately-veined leaves = main vein called midrib with secondary veins branching from it (e.g., elm). – Palmately-veined leaves = veins radiate out of base of blade (e.g., maple). • Parallel venation = characteristics of many monocots (e.g., grasses, cereal grains); veins are parallel to one another. • Dichotomous venation = no midrib or large veins; rather individual veins have a tendency to fork evenly from the base of the blade to the opposite margin, creating a fan-shaped leaf 2. Number of Blades Simple Leaf – blade of single piece. Compound Leaf – blade is composed of two or more separate part on a common petiole. the division is called – Leaflets/Pinnae stalk – petiolules rachis – extension of petiole above the lowest leaflet to upper parts. Stipule-like – stipels Peltate leaves = petioles that are attached to the middle of the blade; examples include mayapple Perfoliate leaves = sessile leaves that surround and are pierced by stems; examples include yellowwort and thoroughwort Mayapple Yellow Wort 3. Compound Leaf classification: Number of leaflets: Unifoliate – 1 leaflet Bifoliate – 2 Trifoliate – 3 multifoliate – more than 3 4. Position of leaflets on rachis Palmately Compound – diverse leaflets from a common point at tip of petiole. Pinnately Compound – petiole becomes into rachis along several leaflets arise at interval. 5. Phyllotaxy or leaf arrangement: Alternate – one leaf to a node of opposite side of stem. Opposite – two leaves to one node of opposite side of stem. Verticulate or Whorld – more than two leaves in one node arranged regularly around the stem. Structures of the Leaf Cuticle – the outermost layer of both the upper and lower surfaces of the leaf. It is clear and waxy to prevent against water loss. Epidermis – a layer of cells one cell thick that provides protection for the inner tissues. These cells are clear to allow light to reach the photosynthetic tissues. Mesophyll – between the epidermal layers. It contains palisade cells that are tall, tightly packed, and filled with chloroplasts for photosynthesis. It also has spongy cells which are irregularly shaped, have large air spaces between them, and fewer chloroplasts. Structures of the Leaf Stomates – openings in the surface of the leaf and stems for gas exchange. The lower surface of a leaf usually has more. Water vapor also passes out through these holes. Guard cells – two of these special cells surround each stomate and regulate the opening and closing of the stomate. Veins – contain the vascular tissue that is continuous with that in the stem. Xylem carries water and minerals upward. Phloem carries dissolved food throughout the plant. INTERNAL STRUCTURE OF LEAVES chloroplasts Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings Conserves water Photosynthesis Transports water and sugar to stem and roots Typical Dicot Leaf Cross-Section Cuticle Palisade Parenchyma Epidermis Vascular bundles Guard Cells Spongy Parenchyma Stoma Typical Monocot Leaf Cross-Section Midvein Vein Bundle sheath cell Epidermis Phloem Xylem Bulliform Cells Stoma Specialized or Modified Leaves • Drought-resistant leaves = thick, sunken stomata, often reduced in size • In American cacti and African euphorbs, leaves are often reduced such that they serve as spine to discourage herbivory and reduce water loss • The stems serve as the primary organ of photosynthesis. Specialized or Modified Leaves • In pine trees, the leaves are adapted to living in a dry environment too. • Water is locked up as ice during significant portions of the year and therefore not available to the plant; pine leaves possess – sunken stomata, – thick cuticles – needle-like leaves – hypodermis, which is an extra cells just underneath the epidermis – Cotyledons or “seed leaves” First leaves produced by a germinating seed Often contain a store of food (obtained from the endosperm) to help the seedling become established. Tendrils Garden Pea Tendrils - blade of leaves or leaflets are reduced in size, allows plant to cling to other objects (e.g., sweet pea and garden peas. INSECT Trapping Leaves Figure 11.8 (1) • Some plants obtain nitrogen from digesting animals (mostly insects). • The Pitcher plant has digestive enzymes at the bottom of the trap • This is a “passive trap” Insects fall in and can not get out • Pitcher plants have specialized vascular network to tame the amino acids from the digested insects to the rest of the plant Specialized Leaves Figure 11.12 (2) • The Venus fly trap has an “active trap” • Good control over turgor pressure in each plant cell. • When the trap is sprung, ion channels open and water moves rapidly out of the cells. • Turgor drops and the leaves slam shut • Digestive enzymes take over Prickles and thorns: epidermal outgrowths on stems and leaves (e.g., holly, rose, and raspberries; Hypodermic trichomes on stinging nettles. Storage leaves or succulent leaves retain water in large vacuoles Bracts: petal-like leaves. Window Leaves: plant is buried in soil with transparent part exposed to light. Being buried reduces loss of war in arid environments. Reproductive leaves, (e.g., Kalanchöe plantlets arise on margins of leaves. Application 1. Identify the parts of the leaf (Petiole, Lamina, Stipules, Midrib, Veinlets) 2. Venation (Parallel, Netted, Obsolete) (pinnately or palmately parallel, dichotomous) (peltate, perfoliate) 3. Compound leaf (leaflets, rachis, petiolules, stipels) 4. Number of leaflets 5. Position of leaflets for compound leaf(pinnately, palmately) 6. Phyllotaxy ( alternate, opposite, whorld) 7. Specialized function (if it has) 8. Blade forms (Linear Lanceolate Oblong Elliptic Ovate Cordate Reniform Spatulate Orbicular 9. Margin ( entire, toothed, dentate, serrate, crenate, incised, lobed, sinnate, undulate)