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EVOLUTION! Summing it all up! What is Scientific Theory? •In science, theories are statements or models that have been tested and confirmed many times. In science, the term "Theory" does not express doubt. •They explain a wide variety of data and observations They can be used to make predictions They are not absolute, can be changed as new evidence is found • • The theory of evolution is considered a UNIFYING THEORY of Biology, because it answers many questions and offers an explanation for observations History of Evolution •Charles Darwin developed the THEORY OF EVOLUTION BY NATURAL SELECTION •explained how organisms changed over time (ADAPTED) Acquired Characteristics •Lamarke's Theory of Acquired Characteristics Some thought that you would gain or lose features if you overused or didn't use them, PROVEN TO BE WRONG! • • Macroevolution: Evolution of a New Species Levels of Evolutionary Study • Microevolution: examines changes to the genes (alleles) within populations – Population Genetics: studies the changes in the numbers & types of alleles in a population • Examines evolution within a species. • Small changes that do not lead to new species, but can lead to new variations. • Macroevolution: examines the evolutionary changes that create new species – Speciation: the formation of new species over time Macroevolution • Species: a group of organisms that can interbreed and produce fertile offspring • Speciation: the evolution of new speices (by genetic change or change in form) –Occurs when members of a similar populations can no longer interbreed! Methods of Speciation • Geographic Isolation: physical barriers divide a population and prevent interbreeding Beetles in Grassland Become Green Beetles in Woody Habitat Become Brown Methods of Speciation • Reproductive Isolation--occurs when formerly interbreeding organisms can no longer mate & produce fertile offspring – Genetic: genetic differences are too great to breed successfully – Behavioral: mating behaviors or mating seasons don’t allow for mating • Chromosome Numbers – Polyploidy: any species with extra sets of chromosomes. Common in plants. • Can interbreed or self-fertilize, forming separate species – Hybridization: when two closely related species attempt to mate • Usually results in sterile offspring • Ex: donkey + horse = mule Speed of Speciation • Gradualism – Idea that species originate thru a slow, gradual change of adaptations over long periods of time • Punctuated Equilibrium – Idea that species can remain stable for long periods of time until environmental changes cause many new species to appear • Speciation occurs in less than 10,000 years! Patterns of Macroevolution: Divergent Evolution • Species diverge or become increasingly distinct from one ancestral species • Ex: Darwin’s finches • Also called adaptive radiation Patterns of Macroevolution: Convergent Evolution • Process by which unrelated species become similar as they adapt to similar environments Ostrich Emu Australia Africa Rhea South America Patterns of Macroevolution: Coevolution • When species who live in close contact evolve adaptations to one another’s existence. Extinction! • When all the members of a species die off or fail to reproduce – Estimated that 99% of all species that ever lived have become extinct – Most extinctions are the result of environmental change, can cause well adapted species to become poorly adapted! – If the environment changes faster than a species can adapt, extinction can result! Microevolution Processes of Microevolution • Microevolution examines the processes by which inherited traits change over time in a population –Natural Selection –Migration –Mate Choice –Mutation –Genetic Drift Processes of Microevolution • Natural Selection: changes in environmental pressures can cause an increase or decrease in certain alleles (traits) in a population – Favorable alleles stay in population (selected for) – Unfavorable alleles are eliminated (selected against) • Migration: as organisms move from one population to another, their genes move with them – Causes the numbers and types of alleles within each population to change. Processes of Microevolution • Mate Choice: if parents are selective (picky) or limited in their choice of mates, only a limited set of traits will be passed on • Mutation: can add a new allele to a population – Mutations are rare, and must occur in egg or sperm to be passed on • Genetic Drift: population sizes affect the change in alleles (traits) in a population – Large populations experience slower changes in alleles – Smaller populations can change more quickly Microevolution: Changes within Populations Populations evolve NOT individuals evolve Natural selection works on the phenotypes within a population Individuals cannot evolve within their lifetime, because they cannot evolve a new phenotype due to a change in the environment Evolution is brought about by breeding that occurs between organisms within a population. Natural selection maintains favorable (useful) genotypes, and eliminates unfavorable (not useful) genotypes. Natural Selection - Explains how organisms change over time 1. Over Production: Most organisms produce more offspring than will survive 2. Variation: Members of the population have variations 3. Selection: Certain variations will increase the likelihood of survival 4. Adaptation: Over-time organisms with that variation make up most of the population and may look different than their ancestors Patterns of Natural Selection Stabilizing selection: Directional selection: Favors average individuals; reduces variation Favors EXTREME variations; evolution is fast Disruptive selection: Favors two extreme variations; no intermediate and eventually two species. Stabilizing Selection • “Average” individuals are better able to survive • Alleles for the “extreme” are eliminated • A narrow range of phenotypes is favored resulting in a narrow bell shaped curve Directional Selection • ONE extreme is favorable, but other is not • Alleles for other extreme and normal phenotypes become less common in the population • Results in FAST evolution! • Bell curve is shifted off of center Disruptive Selection • BOTH extremes are favorable! • Results in increasingly distinct phenotypes, the “normal” phenotype is selected against • Bell curve is ‘disrupted’ and pushed apart into two peaks Evidence for Evolution Evidence for Evolution • Fossils – provide a record of early life & evolutionary history – Currently, the fossil record is incomplete (like a jigsaw puzzle w/ missing pieces) – The more fossils discovered, the clearer the evolutionary picture becomes! Evidence for Evolution • Biogeography –The study of the locations of organisms around the world –Organisms in similar environments have similar adaptations despite being on completely different continents. •Ex: Large, flightless birds Evidence for Evolution • Anatomy –the way an organism is put together –Homologous structures –Analogous Structures –Vestigial Structures Homologous Structures • Parts with a common evolutionary history – Ex: Organisms that have the same basic skeleton, bones are just modified for different functions – It would be unlikely for so many animals to have similar structures if each species arose individually! Analogous Structures • Body parts that are similar in function, but not evolutionary origin • Ex: Wings of a butterfly & wings of a bird Vestigial Structures • Body structure that has no function in present day organisms, but was probably useful to an ancestor • Ex: Eyes on Naked Mole Rats, Appendix in humans, Wings on ostrich Evidence for Evolution • Adaptations: Structural – Develop over many generations (ex: claw length, fur color, etc) •Mimicry: enables one species to resemble another species. Monarchs and Viceroys •Camouflage: enables species to blend in with their surroundings Monarch TOXIC! Viceroy TASTY! Evidence for Evolution • Adaptations: Physiological – Changes in an organisms metabolic processes (occurs more rapidly) • Bacteria’s resistance to antibiotics • Insects / weeds resistance to certain pesticides Evidence for Evolution • Embryological – similarities between young embryos suggest evolution from a distant common ancestor Evidence for Evolution • Biochemical – Provides info about relationships between individuals and species – Uses RNA & DNA sequences to construct cladograms (evolutionary diagrams)