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Chapter 15 Our Senses Vision Smell Taste Hearing Balance Touch* Anatomy of an Eyeball Accessory structures 3 tunics (layers) Fibrous (blue) Vascular (yellow) Sensory (red) Segments Anterior divided into chambers Posterior Filled with humors (fluid) Lens Accessory Eye Structures Eye muscles Rectus as named; obliques lateral and opposite Diplipia: muscle weakness/alcohol Strabismus: eyes not aligned Eyebrows Eyelids Blink to spread secretions Eyelashes trigger blinking Conjunctiva Mucus prevents drying out Conjunctivitis Lacrimal apparatus Tears clean, protect, and moisten Excess secretions Emotional tears unique to humans Stuffy/runny nose when cry Watery eyes with cold Fibrous Tunic Avascular CT Sclera (white of the eye) Protects and shapes Muscle attachment Continuous with dura mater Cornea (transparent) Outer stratified squamous, why? Inner simple squamous maintain clarity Innervated Transplants not rejected between people Vascular Tunic Choroid Vascularized to supply nutrients Melanocytes to absorb light Ciliary body Smooth muscle ring ciliary muscles control lens shape Ciliary processes secrete aqueous humor Suspensory ligaments hold lens in place Iris Colored portion of ciliary body Brown pigment only (varies) Less scatters light = blues/greens/grays (babys) Encircles the pupil (2 smooth muscle layers) Sensory Tunic Pigmented layer (outer) Prevents light scattering Phagocytize damaged photoreceptors Neural layer (inner) Photoreceptors, bipolar cells, ganglion cells Rods and cones Blind spot (optic disc) filled Macula lutea and fovea centralis Rapid eye movement for rapid scene changes Vascular supply from choroid and central vein/artery Opthalmologist examines Retinal detachment when layers separate Vitreous humor seeps in Photoreceptors lose nutrients = blindness Humors Anterior segment with aqueous humor Similar to blood Continual development Nutrients & O2 to lens, cornea, & retina Blocked drainage = up pressure = glaucoma Posterior segment with vitreous humor Transmits light, support lens, & intraocular pressure Embryonic development The Functioning Eye Light enters the pupil, regulated by the iris Passes through a convex lens Avascular Lens fibers added through life Cataracts = clouding of lens due to loss of nutrients Lens is shaped by the ciliary body to focus light on the retina (accommodation) Refraction of light converges to a focal point Real image forms upside down and reversed Visual Pathways Visual field Overlap to provide depth perception = 3D vision Ganglion cells Optic nerve Optic chiasm Nasal and temporal visual field Optic tract Thalamus LGN Primary visual cortex Conscious perception of images Olfactory Receptors Ciliated bipolar cells Located in olfactory epithelium (psuedostratified ) Mucus captures and dissolves odorants Pass through cribriform plates Synapse in olfactory bulbs Odorant detection Humans can distinguish 10,000 odors Some is pain (ammonia, chili, methanol Combinations of different odorant/receptor binding Olfactory Neural Pathway Olfactory receptors synapse with mitral cells Contained in glomeruli Receptor type specific Refines smell Mitral cells signal via olfactory tracts 2 pathways Olfactory cortex Hypothalmus, limbic system = emotional connection Taste Taste buds detect molecules in solution About 10,000 Four familiar and 1 other found in papillae Sweet: organic substances Alcohol, sugar, amino acids Sour : acids, H+ in solution Salty: inorganic salts Bitter: alkaloids Aspirin, nicotine, caffeine Umami: glutamate & aspartate meats, cheeses, and protein-rich foods (MSG) Each receptor responsive to a particular type of substance Often mixes Many ‘tastes’ (80%) are really smell (head colds) Papillae Fungiform Mushroom shaped Tops of, all over tongue Foliate Fold is side walls Circumvallate Largets, fewest, back of tongue Filiform Hair like projections all over tongue Do not have taste buds Gustatory Neural Pathway Cranial nerves carry sensations to medulla Relay through the thalamus into primary gustatory cortex Pathway initiates digestive process too Regions of the Ear Outer ear Pinna, external auditory canal, and tympanic membrane (separates) Middle ear Pharyngotympanic tube equalizes pressure b/w middle ear and atmosphere (‘pop’) Function of tympanic membrane Ossicles (malleus, incus, & stapes) amplify signal Inner ear Bony and membranous labyrinths Semicircular canals for rotation of head Vestibule report on changes of head position Saccule and utricle with equilibrium sensors called maculae Cochlea houses the hearing organ The Cochlea Scala vestibuli Perilymph: like CSF Oval window Scala Tympani Perilymph Round window Scala media (Cochlear duct) Endolymph: K+ rich intercellular fluid Organ of Corti Contains hair cells embedded in a basilar membrane Vestibular membrane Tectorial membrane bends cells as basilar membrane moves Signal to auditory nerve Frequency and Amplitude Pitch depends on frequency High pitch = higher frequency Basilar membrane responsive to certain frequencies Sounds detected as increase in AP’s by the brain Greater volume = higher amp of generated wave Vigorous vibrations in cochlea = more bending = more AP’s Hair cells easily damaged due to prolonged exposure to certain frequencies Physiology of Hearing Pinna collects sound waves Travel down auditory canal to tympanic membrane Moves ossicles as it vibrates Stapes pushes on oval window, in and out Creates fluid pressure waves in scala vestibuli perilymph Pressure waves deform scala tympani to push round window in and out Pressure changes move endolymph Pressure changes in endolymph, from perilymph changes, moves the basilar membrane Hair cells on Organ of Corti bend as they move against the tectorial membrane Generates nerve impulses that leave via the cochlear nerve Auditory Pathway AP signals from cochlea to medulla Cochlear nuclei Some fibers cross, all ascend, from olives into MGN in the thalamus Pass through inferior colliculi (reflex area) Interactions with superior colliculi to turn toward sound Synapse in primary auditory cortex Localization utilizes relative intensity and timing http://openlearn.open.ac.uk/file.php/3373/SD329_1_027i.jpg Equilibrium Static Respond to changes only, keeps head still and balanced Maculae embedded in otolithic membranes in saccule/utricle Dynamic Maintenance of body position after sudden movement Crista in ampullae of semicircular canals Both operate by bending hair cells Head movements cause fluid movement (slower) Changes action potentials Sudden stop, but fluid still moving = dizzy Brain interprets and commands skeletal muscles Motion Sickness Results from conflict between eyes and equilibrium sensors in the inner ear Feeling motion, but not seeing it (inside car) One system is hallucinating, implying toxins in system = vomiting Dramamine inhibits input from equilibrium sensors Astronauts learn to control