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March 28, 2005 Plant Tissues Chapter 26 Jin Hoe Huh Angiosperms – flowering plants • The angiosperms are seed-bearing vascular plants • In terms of distribution and diversity, they are the most successful plants on Earth • The structure and function of this plant group help explain its success Flowering Plant Life Cycle Diploid Double fertilization Haploid pollination Two sperms enter ovule Meiosis microspores Female gametophyte Meiosis Mitosis without cytoplasmic division Plant Life Histories • Annuals complete life cycle in one growing season • Biennials live for two seasons; flowers form in second season • Perennials grow and produce seeds year after year Shoot and Root Systems Shoot system - produces sugars by photosynthesis Shoot System - carries out reproduction Root system - anchors the plant - penetrates the soil and absorbs water and minerals - stores food Root System Shoot and root systems are interdependent water & minerals sugar SHOOT SYSTEM ROOT SYSTEM Plant Tissue Systems • Ground tissue system • Vascular tissue system • Dermal tissue system EPIDERMIS VASCULAR TISSUES GROUND TISSUES SHOOT SYSTEM ROOT SYSTEM Meristems – Where Tissues Originate • Regions where cell divisions produce plant growth • Apical meristems – Lengthen stems and roots – Responsible for primary growth • Lateral meristems – Increase width of stems – Responsible for secondary growth Apical Meristems Lengthen shoots and roots: SAM and RAM activity at meristems Cells that form at apical meristems: protoderm epidermis new cells elongate and start to differentiate into primary tissues ground meristem ground tissues procambium primary vascular tissues Lateral Meristems Increases girth of older roots and stems Cylindrical arrays of cells vascular cambium secondary vascular tissues periderm cork cambium thickening Simple Tissues Made up of only one type of cell Parenchyma Collenchyma Sclerenchyma Morphology of three simple tissue types parenchyma collenchyma sclerenchyma Parenchyma: A Simple Tissue • Comprises most of a plant’s soft primary growth • Cells are pliable, thin walled, many sided • Cells remain alive at maturity and retain capacity to divide • Mesophyll is a type of parenchyma that contains chloroplasts Collenchyma: A Simple Tissue • Specialized for support for primary tissues • Cells are elongated, with walls (especially corners) thickened with pectin • Makes stems strong but pliable • Cells are alive at maturity Sclerenchyma: A Simple Tissue • Supports mature plant parts • Protects many seeds • Cells have thick, lignified walls and are dead at maturity • Two types: – Fibers: Long, tapered cells – Sclereids: Stubbier cells Complex Tissues Composed of a mix of cell types Xylem Phloem Epidermis Xylem • Conducts water and dissolved minerals • Conducting cells are dead and hollow at maturity tracheids vessel member Phloem: A Complex Vascular Tissue sieve plate • Transports sugars • Main conducting cells are sievetube members • Companion cells assist in the loading of sugars sieve-tube member companion cell Epidermis: A Complex Plant Tissue - Covers and protects plant surfaces - Secretes a waxy, waterproof cuticle - In plants with secondary growth, periderm replaces epidermis Monocots and Dicots – same tissues, different features 1 cotyledon 4 or 5 floral parts 3 floral parts Parallel veins 1 pore Vascular bundles in ring 2 cotyledons Netlike veins 3 pores Vascular bundles dispersed shoot apical meristem Shoot Development cortex procambrium protoderm procambrium pith ground meristem primary xylem primary phloem Bud = undeveloped shoot of meristematic tissue Leaves Internode Axillary bud at node Longitudinal section of terminal bud Roots also have meristems Internal Structure of a Dicot Stem - Outermost layer is epidermis - Cortex lies beneath epidermis - Ring of vascular bundles separates the cortex from the pith - The pith lies in the center of the stem Internal Structure of a Monocot Stem • The vascular bundles are distributed throughout the ground tissue • No division of ground tissue into cortex and pith Dicots Monocots Ground tissue system Dermal tissue system Vascular tissue system Dicots and Monocots have different stem and root anatomies Leaf Gross Structure DICOT MONOCOT petiole axillary bud blade node sheath blade node Adapted for Photosynthesis • Leaves are usually thin – High surface area-to-volume ratio – Promotes diffusion of carbon dioxide in, oxygen out • Leaves are arranged to capture sunlight – Are held perpendicular to rays of sun – Arrange so they don’t shade one another Leaf Structure UPPER EPIDERMIS cuticle PALISADE MESOPHYLL xylem SPONGY MESOPHYLL phloem LOWER EPIDERMIS O2 CO2 one stoma Mesophyll: Photosynthetic Tissue • A type of parenchyma tissue • Cells have chloroplasts • Two layers in dicots – Palisade mesophyll – Spongy mesophyll Collenchyma Parenchyma Leaf Veins: Vascular Bundles • Xylem and phloem – often strengthened with fibers • In dicots, veins are netlike • In monocots, they are parallel Root Systems Root Structure • Root cap covers tip • Apical meristem produces the cap • Cell divisions at the apical meristem cause the root to lengthen • Farther up, cells differentiate and mature root apical meristem root cap Internal Structure of a Root • Outermost layer is epidermis • Root cortex is beneath the epidermis • Endodermis, then pericycle surround the vascular cylinder • In some plants, there is a central pith epidermis endodermis cortex pericycle root hair phloem xylem Root Hairs and Lateral Roots • Both increase the surface area of a root system • Root hairs are tiny extensions of epidermal cells • Lateral roots arise from the pericycle and must push through the cortex and epidermis to reach the soil new lateral root Secondary Growth • Occurs in perennials • A ring of vascular cambium produces secondary xylem and phloem • Wood is the accumulation of these secondary tissues, especially xylem Secondary Growth Woody Stem periderm (consists of cork, cork cambium, and secondary cortex) BARK vascular cambium secondary phloem HEARTWOOD SAPWOOD Annual Rings • Concentric rings of secondary xylem • Alternating bands of early and late wood • Early wood – Xylem cells with large diameter, thin walls • Late wood – Xylem cells with smaller diameter, thicker walls Types of Wood • Hardwood (oak, hickory) – Dicot wood – Xylem composed of vessels, tracheids, and fibers • Softwood (pine, redwood) – Gymnosperm wood – Xylem composed mostly of tracheids – Grows more quickly