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Adaptation • Adaptation – describes any trait that enhances an organisms fitness or increases its chance of survival. • While adaptations are products of natural selection, variations within a species are the raw material upon which natural selection acts. • Exaptation – an adaptation that evolved for one function that could also be used for another purpose. Limbs on a lungfish Types of Adaptations • 1. Structural (anatomical) Adaptations – these may include external or internal changes within an individual, or even mimicry and cryptic colors can be considered physical. • 2. Physiological adaptations – changes within the function of the organism. – Enzymes – Chemical defenses • 3. Behavioral adaptations – how an organism responds to the environment. – Migration – courtship • Is evolution perfect???????? Speciation • WHAT IS SPECIATION? – The formation of a species (a reproductively compatible population , ie. a population that can interbreed and produce viable offspring) • There are two pathways that lead to new species: transformation vs. divergence pg 708 – For species to remain distinct, they must remain reproductively isolated, or prevent breeding between closely related species lines. How to accomplish speciation? • GEOGRAPHIC BARRIERS or ISOLATION • REPRODUCTIVE (Biological ) BARRIERS or ISOLATION http://evolution.berkeley.edu/evolibrary/home.php Life Sciences-HHMI Outreach. Copyright 2006 President and Fellows of Harvard College. Geographic Isolation • Nature has frequently provided geographical barriers that prevent individuals from interbreeding. • Given sufficient time and generations new species may emerge. Biological Barriers • Even when individuals live in the same region factors may exist that prevent interbreeding: – Pre-zygotic Barriers – impede the mating process or prevent the egg from becoming fertilized – Post-zygotic Barriers – if sperm from one species is capable of fertilizing the egg of another species. There are mechanisms to prevent this hybrid individual from developing. Types of Pre-Zygotic Barriers • Behavioural Isolation – a species may have special signals or rituals that is species specific. – Eg. Songs of birds, pheromones etc… • Habitat Isolation – two species can live the same region but occupy different habitats. – Eg. Garter snake lives near water where most other snakes prefers open areas. Pre- Cont. • Temporal Isolation- species may be isolated due to the time or season they are active or are reproductively viable. • Mechanical Isolation- related species may attempt to mate but are anatomically incompatible. Pre- Cont. • Gametic Isolation – even if gametes from different species do meet, rarely will a zygote form. – Sperm from one species can not survive in the female reproductive tract of another species. Post- Zygotic Barriers • Hybrid Inviability – development of the hybrid zygote is stopped at some stage during embryonic development. • Hybrid Sterility – when two different species can mate but the offspring is sterile. – Horse + Donkey = Mule (sterile) • Hybrid Breakdown – when first generation hybrids are viable and fertile but if these hybrids are mated, the next generation are sterile or weak. Again Not Perfect • The biological definition of a species does not work in all instances… (discuss) • Other models include: – Morphological species concept (physical or phenotype characteristics) – Cohesion species concept – Ecological species concept – Evolutionary species concept PATTERNS OF SPECIATION? How does it occur? Two Modes Of Speication: (Pg. 714) 1. Sympatric Speciation: when reproductive isolation happens when species are not geographically isolated. 2. Allopatric Speciation (geographical speciation): when speciation happens in species that are geographically separated. • GRADUALISM • PUNCTUATED EQUILIBRIUM Life Sciences-HHMI Outreach. Copyright 2006 President and Fellows of Harvard College. Adaptive Radiation • The creation of new species that have radiated from a common ancestral species that have adapted differently to their various environments. • DIVERGENT EVOLUTION and CONVERGENT EVOLUTION Life Sciences-HHMI Outreach. Copyright 2006 President and Fellows of Harvard College. Divergent Evolution • The Pattern of evolution where species that were once similar to an ancestral species diverge, or become increasingly distinct. ADAPTIVE RADIATION – AN EXAMPLE OF DIVERGENT EVOLUTION http://www.vanderbilt.edu/AnS/english/Clayton/Galapago_finches.gif http://evolution.berkeley.edu/evolibrary/home.php Life Sciences-HHMI Outreach. Copyright 2006 President and Fellows of Harvard College. Convergent Evolution • Similar traits arise independently in species to adapt and survive in similar environmental conditions. They DO NOT share a common ancestor. CONVERGENT EVOLUTION – WHAT IS AN EXAMPLE? http://evolution.berkeley.edu/evolibrary/home.php Life Sciences-HHMI Outreach. Copyright 2006 President and Fellows of Harvard College. Comparing the anatomy of organisms indicates a common ancestry because of: • homologous structures - structures having a common ancestry but with different uses in various species. – Eg. Similar bone structure of the forelimb of a bat, whale, horse and human suggests these different species have a similar evolutionary origin. Page 113,114 & 664 • - analogous structures - body parts of organisms that do not have a common evolutionary origin but perform similar functions. –Eg. insect wings and bird wings are similar in function but not in structure. Page 665 - vestigial organs - small or incomplete organs ( or bones ) that have no apparent function in one organism but do have a function in another species. This indicates evolutionary origin from a common ancestor. Page 665 – Eg. Human ear muscles, Human appendix, Hip bones in whales, Human tail bone, Leg bones in snakes, and Forelimbs in the flightless ostrich Co-Evolution – the evolution of species together so that both find ways to adapt and survive.