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Chapter 17 Speciation and Molecular Evolution Reproductive Isolation • Species: a group of organisms that can mate with each other an produce fertile offspring • Speciation: the formation of a new species • Reproductive Isolation – when members of a population stop reproducing with each other and this eventually leads to evolution into two separate species – Mechanisms for reproductive isolation: behavioral, geographic, and temporal Behavioral isolation • Use of different courtship rituals or behaviors that prevent related organisms from reproducing • Western meadowlark song: http://www.yo • utube.com/watch?v=lvAUgFb1cLY • Eastern meadowlark song: • https://www.birdjam.com/birdsong.php?id=4 3 Geographic isolation and temporal isolation • Geographic isolation: Separation of populations by rivers, mountains, or water • Temporal isolation: timing of reproduction isn’t in sync – Ex. Timing of flowering plants Hypothesizing about Speciation in Darwin’s Finches • Founding of a new population (founder effect) – original species on island reproduced, change in allele frequency compared to population in S. America • Geographic isolation – movement to different islands caused isolation and speciation • Changes in the gene pool – adaptation to local environments on each island • Behavioral isolation – movement back to original island – no reproduction between these species • Ecological competition – more evolution due to natural selection Molecular Evolution • Molecular clock: rates of DNA mutations used for estimating the time that two species have been evolving independently • Analyzing Data: Fishes in Two Lakes page 500