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Transcript
Chapter 6 Seeding Methods Successful revegetation(restoration, rehabilitation, or reclamation) is obviously dependent on the germination, establishment and eventual reproduction of desired seeded species. All reasonable efforts effort must be made to assure suitable seed to soil contact at the most appropriate time of year. All planning processes may have gone smoothly and intelligently up to this point, but if the seed being broadcast (or drilled or hydroseeded) on the ground is not done in a proper manner, all is for not. This is where the rubber really starts meeting the road- so to speak or where the seed starts meeting the soil. Although seeding is generally the most widely used method for restoration and revegetation, planting live plants is also an option for smaller projects. Although planting live plants is more expensive, it may be the best option in certain situations and may result in more rapid establishment. Some projects may use a combination of planting and seeding for desired results. It should also be noted that natural revegetation should also be considered as it may be the best option when both native plants are already established in a reasonable density near the site and when weeds do not seriously threaten to outcompete the native plants. It is important to consider not only the patterns of the natural landscape for designing planting patterns, but to also consider different species abilities to respond to different soil types, aspects, slopes, etc to assure seeded species are placed in appropriate locations. Sometimes different seed mixes and/or seeding methods for the same restoration site are warranted to best mimic the natural landscape. It must be noted that seeding native species often requires experience, but always requires patience. Expect to wait at least three years before seeing significant results of a seeding project, but often, five years are necessary. SEEDING The most common method of restoration is the use of specialized seed mixes. As mentioned in Chapter 2, the underlying characteristics of each site (soils, aspect, slope) should be carefully considered prior to determining seeding methods as well as seed mixes. The numerous vegetation cover types of the Colorado Plateau highlights the variability of potential site conditions and thus prevents the use of only one set of seeding guidelines. However, it should be noted that plant establishment is more likely to be successful if locally adapted species are used, particularly if properly seeded at the correct time. Prior to seeding, pose the following questions once again to assist proper planning prior to seeding and project ‘after-care’: Is seeding necessary? Do undesirable plants dominate the site? Is weed control practical? Is the desired amount and type of seed available and affordable? Should additional preparation be done before planting is undertaken? Answers to these questions will help in the determination of the feasibility, timing, cost, and probability of success of the project. TIME OF SEEDING As many restoration projects do not involve irrigation, the planting window must be chosen to take advantage of natural moisture. Higher seeding success is directly proportional to temperatures above freezing and high precipitation (provided it does not all come at once!). On the Colorado Plateau, the best seeding window for cool season species is usually the early spring and late fall. It is critical to truly wait until the late fall for seeding as many projects have failed as a result of seedling emergence in the fall, but then perishing in an early frost. For warm season species, seeding in late spring or early summer is better as temperatures are more likely to be warm enough to support germination. Seeding should not be done when the ground is frozen. It is important to seed a site as soon as final grading and topsoil placement have occurred so as to minimize erosion and weed establishment. If seeding cannot be accomplished in a timely manner, bare soils should not remain unprotected. In these cases, it is possible to sow a temporary cover crop such as ReGreen or other sterile hybrids to provide some soil cover until the final seeding can be accomplished. SEEDING RATES Often, seeding rates are expressed in terms of pounds per acre. This can often be misleading as the number of seeds per pound varies greatly by species. For instance, sand dropseed (Sporobolus cryptandrus) has more than 5 million seeds per pound, whereas bluebunch wheatgrass (Pseudoroegneria spicata) has only 140,000. If only pounds per acre are specified, the final seed mix can easily become 1 pound each of sand dropseed and bluebunch wheatgrass among some other possible species. The final seed mix would equate to 98% sand dropseed and 2% bluebunch wheatgrass and would put 249 seeds per square foot on the ground – a rate that is much too high and would be wasteful of valuable seed. It is much more precise to create seed mixes that concentrate on the number of seeds per square foot. The seed mix calculator available XX(WHERE) concentrates on assuring the correct percentages of each species desired in the mix by taking into account the seed size or number of seeds per pound. Many opinions exist as to the ideal number of seeds per square foot for a successful restoration project. However, the ideal number will vary with project goals as well as site conditions. The following are suggested seeding rates for common site conditions and seeding methods. Seeding Method/ Site Conditions Broadcast Hydroseeding Drilling Bare ground 60- 80 seeds/sq ft 60-80 seeds/ sq ft 30-50 seeds/ sq ft Inter-seeding /over seeding 20-40 seeds/ sq ft 20-40 seeds/ sq ft ~20 seeds/ sq ft SEEDING METHODS The three primary seeding methods include drilling, broadcasting and hydroseeding. The best method to use will depends on site accessibility and terrain, seedbed characteristics and time of seeding. Furthermore, each seeding method has an associated ideal seeding rate as well as technique, given the site’s characteristics. DRILL SEEDING Drill seeding is used most often as it has generally shown the highest success rate. This is generally because of the high seed to soil contact achieved through drilling as well as the ability to better control seeding depth with some specialized drill seeders. However, drill seeders tend to lose some effectiveness on slopes steeper than 3:1, where much standing woody materials (e.g. juniper or pinyon) occurs, or where the soils are extremely rocky. As with any seeding method, there are benefits and liabilities to drill seeding. Benefits include: Generally higher seeding success Ability to better control seeding depth TIPS FOR DRILL SEEDING In general seeding to a depth of 1/4 to 1/2 inches should be adequate. Seeding depth recommendations vary from species to species and are included in Appendix (?). When seeding multiple species, one can use a rangeland drill, such as the Truax drill (see photo below), which has the capability of seeding different species at different depths at the same time. Seeding should be conducted along the contour to avoid erosion from water flowing down drill furrows. Truax rangeland Drill (photo from http://www.aprairiehaven.com/) If a Truax drill is not available, greater success can often be achieved with smaller grass seed and forbs by placing them in a separate seed box and alternating the seeing depth between rows or dropping them directly on the ground to be covered by the action of heavy, trailing chains. **ALWAYS check seeding depth in a test strip prior to seeding the entire project! The following components are required in a drill suitable for general use in native seeding (Figure 7.1): Multiple seed boxes for different types of seed (agitators and picker wheels in at least one box for fluffy seed). Double disc furrow openers (to cut through residue and reduce potential clogging problems. The double-disk opener has good depth control under conservation tillage conditions because it does not have a tendency to float up and over residue) Intact depth bands with functioning scrapers on all disc openers to ensure consistent, uniform seed depth placement Seed tubes, which drop between disc openers, large enough to handle fluffy seed Packer wheels with adjustable tension, to provide proper soil compaction over and adjacent to the seed Coulter wheels to allow penetration of furrow openers where seeding into heavy mulch or cover crop adjacent to the seed Seed Boxes With Agitators Double Disk Furrow Opener Depth Band Scraper Packer Wheels Coulter Wheel Figure 7.1 – Illustration of important components of a drill suitable for general use in native seeding as outlined above. Liabilities of drilling include: Seeds drilled in rows may suffer from high interseedling competition Unless specially modified drills (such as the Truax drill) are used, all seeds, regardless of size, will be planted at the same depth; the smallest seeds are likely to be planted too deep. Drill seeding leaves “rows” which often persist for many years; this may be a visual or aesthetic liability. For more information on variations of seeders, see Chapter 9 of USDA Forest Service Gen. Tech. Rep. RMRS-GTR-136. Restoring Western Ranges and Wildlands, 2004 See Appendix X for basic calibration of drill seeders and notes on calibration of broadcast seeders BROADCAST SEEDING Broadcast seeding is best on difficult terrain (slopes that are steep, extremely rocky, with excessive amounts of standing woody materials, remote or inaccessible locations). For broadcast seeding, it is critical to pay close attention to seedbed preparation, especially being sure to harrow/ rake/ chain (or similar method) both before and after seeding to ensure good seed to soil contact and eliminate any potential soil crusting. It is important to note that: Broadcast seeding requires at least doubling the seeding rate of drill seeding Benefits of broadcast seeding include: The variable planting depths that result from broadcast seeding allows better establishment of smaller seeds than with drill seeding. Resulting vegetation not in “rows”, which is more aesthetically acceptable on many sites. Liabilities of broadcast seeding include: High amount of seed necessary Imprecise calibration of seeding rates If broadcasting is not performed correctly, germination and seedling establishment tends to be slower Broadcast seeding should not be attempted on windy days Careful calibration of seeding rates and methods should be done to assure uniform coverage over the site. HYDROSEEDING A hydroseeder consists of a water tanker with a special pump and agitation device to apply the seed under pressure in a water slurry (Figure 7.2). Seed is then sprayed on to a roughened slope. The results of hydroseeding are generally less satisfactory than drill or broadcast seeding because the seed does not make good soil contact. In addition, the slurry mix often rolls off the steep, hard slopes it is applied to, leaving very erratic and uneven distribution of seed. Hydroseeding first application Applying second layer of mulch More success has been reported by making hydro-seeding a 2-stage process, whereby seed is hydraulically applied with enough mulch to act as a tracer, then the majority of the mulch is applied over the top in a second layer. This process has improved stand establishment, but overall success is still poor relative to drill seeding or broadcasting/harrowing, where seeds are actually incorporated into the soil. In addition, several thousand gallons of water are used to apply seed and mulch over one acre, making hydro-seeding the most expensive seeding method available. Hydroseeding may be a good choice for seed that needs sunlight for germination, such as sand dropseed and sagebrush. However, it should be limited to steep, inaccessible slopes in areas with adequate and dependable moisture during the growing season. However, be aware that the sheet flow of sprayed water on steep, impermeable slopes may wash the seeds off the slope. Alternative Methods for Successful Plantings [Columbia R Basin handbook] As mentioned previously, it is important to revisit restoration objectives and seeding conditions in the planning process. If invasive species pose a serious risk to the success of the project, options available are discussed below to allow for greater success. Staged Plantings Many restoration projects that include forb or shrub seeding have utilized a “staged planting” approach, whereby grasses and forbs are planted in successive years. This approach allows one year of weed control with broadleaf-selective herbicides after grasses have been seeded. There are pros and cons to this approach, but on balance, it seems to be the most effective technique currently available. Weed Control Tips For weed control in these staged plantings, there is a limited time window immediately following seeding when cheatgrass and other weeds can be controlled with no impact to seeded species. The length of this window depends on soil moisture and temperature; germination and emergence of many native grasses will occur within 7 – 10 days with good soil moisture and soil temperatures at or above 45oF. If soil temperatures are between 32oF and 45oF, seeded species will emerge slowly, taking 2 – 3 weeks. Cheatgrass control is critical in the fall and early winter as fall-emerged cheatgrass is very competitive with young grass seedlings. Cheatgrass roots may already be established and depleting soil moisture by the time seeded species germinate and emerge (Hironaka 1961). Prior to seeded species emergence, cheatgrass and other weeds can be controlled with herbicides such as Roundup that have no soil activity. The rate of Roundup used for cheatgrass control depends on the growth stage of cheatgrass and seeded grasses. As the likelihood of planted species emergence increases, Roundup application rates should decline to minimize impacts to seeded species. Seeding with a seed drill or air seeder/harrow typically results in dust deposition on cheatgrass leaves, which may preclude cheatgrass control by reducing herbicide absorption through the leaf (Zou and Messersmith 2005). Dusty leaves are normally only an issue for several days after seeding; rain, wind, or several nights of dew or frost is sufficient to clean dust off of leaves. Should herbicide application occur within several days of seeding, cheatgrass leaves should be examined for dust, and application postponed as needed (no more than a few days) to achieve good herbicide contact. Planted Stubble Mulch Crops Annual grasses, such as sterile forage sorghums, sudan, or forage millets, can be planted the growing season prior to permanent seeding. After crop maturation, native seeds can be sown into the residual standing dead material, as this material serves to protect seedlings from potential harsh winds and provide mulch to retain moisture. This is a very cost effective way to provide mulch and seedling protection. It should be noted that this method differs distinctly from use of a “nurse crop” in which the annual grain and the perennial mix are planted simultaneously. The “nurse” usually ends up outcompeting the slow-growing perennials, especially where moisture is limiting. Wheat, rye, and barley should not be used unless they will be mowed before seed maturity, as they will produce seed that will compete with the seeded native species. The annual forage graminoids must be mown prior to seed maturation to prevent reseeding and allow easier drilling of the permanent seed. Planting into such cover requires a grass or no-till seed drill equipped to handle low tillage and high “trash” conditions. Sterile hybrids (such as ReGreen or Quick Guard) can be used for this purpose and do not have to be mowed prior to planting. Stubble mulch is most appropriate on low-slope sites because water erosion control capabilities of such covers are limited. Forb Islands Sometimes the expensive nature of forb and shrub seed can limit the amount that can be used in a restoration project. One approach to balance costs with habitat objectives is to seed only a portion of the project with these species according to funding and seed availability. Using this approach, forb and shrub seed is planted in strips in a planned pattern (strips or located by most appropriate habitat) to provide the maximum opportunity for seed movement after source plants have become established. Depending upon conditions, the speed of forb and shrub movement into unseeded areas may be relatively fast for wind dispersed species such as yarrow, fleabane daisy, and sagebrush, but significantly slower for species with limited seed dispersal such as penstemons and balsamroot. However, if sown grasses establish well on a site, subsequent spread of forbs and shrubs into this competitive matrix may be extremely limited, at least over the short term. Additional long-term monitoring is needed to fully assess the efficacy of this approach in establishing a diversity of native forbs. Planting plugs Growing seedlings in the greenhouse and out-planting into the field is commonly used in restoration scenarios. Seedlings can be planted in areas where soils are too rocky for traditional site preparation, and/or to accelerate wildlife habitat development (Newsome 2011). Seedlings can also be used to augment plant communities that are somewhat degraded, but still have some components of the native plant community that managers wish to retain. In instances where seed is limited, (e.g. rare plant species), planting seedlings can also make the most efficient use of limited seed. Outplanting of plugs may also be used with species that have shown to establish poorly in the field via seed. When this method is used, it is important to consider irrigation needs. Plants should be watered at the time of planting, and will benefit greatly from supplemental waterings. Generally, plants will require at least one year of regular waterings. A soaking once every three weeks should be sufficient in cool weather, with increased waterings during warm periods. In some instances, packages of Dri-Water (or approved equivalent) can be planted alongside the roots of the plugs to assure some moisture during establishment. Some have planted PVC tubes beside plugs into which Dri-Water can be inserted repeatedly as needed until establishment. Alternatively, a long PVC pipe can be buried next to the plug into which water can be placed to assure the roots receive the moisture they need. Photo of plant with associated deep pipe irrigation tube Photo courtesy of David Bainbridge Fertilizers and/or Root Growth Hormones Available literature differs on its opinions concerning fertilizer use at the time of transplanting. Ostler and Allred (1987) state that Osmocote tablets, a commercial slow-release fertilizer, should be included at the bottom of the transplanting hole to assist with establishment. Some professionals report that many native species, especially those with increased drought tolerance, react adversely to fertilizer use at the time of establishment. With such species, even mild fertilization can cause root-dieback and shoot burning. It is an excellent idea to consult with a nursery, a plant ecologist, or the Natural Resource Conservation Service (NRCS) to help determine specific requirements or suggestions. POST SEEDING After seeding, most sites will benefit from placement of a protective mulch cover. Such covers protect soil and seeds from erosion by wind and water, and conserve soil moisture from the effects of wind and sun. To be effective, mulches must cover the ground nearly completely and have sufficient durability to survive until the seeds germinate. MULCH In general, mulch should be applied immediately after seeding to protect seed and to avoid disturbing germinating seeds. The following are commonly used mulch types: HAY In general, hay mulches are more durable the longer the average unbroken stem length is. Loose hay is usually highly susceptible to being blown off the surface in most Colorado Plateau sites. To counteract this issue, hay should be “crimped” into the soil surface by using a modified disc plow to jam the hay stems into the soil. This does little directly to enhance or stabilize the important cover function of mulch but it can mimic the effects of reducing surface wind speeds and soil desiccation that stubble mulch provides. Durability of the applied mulch cover is also enhanced by addition of organic tackifier products that “glue” (at least temporarily) the hay to itself and to the ground. When used, hay is typically applied at the rate of 3,000 to 4,000 pounds per acre, but sites can benefit from rates as low as 2000 pounds per acre. Depending upon the hay market, the benefit of applying a mulch far outweighs the initial cost. However, hay may introduce undesirable weeds or nonnatives (e.g. smooth brome hay) that may jeopardize carefully planned restoration projects. Beginning November 1998, users of BLM administered land in Utah will be required to use only certified noxious weed free hay, straw or mulch. Approved products for livestock feed on public lands include pellets, hay cubes, processed grains and certified hay, straw or mulch normally available at some feed stores and producers in Utah. (http://www.blm.gov/ut/st/en/fo/vernal/grazing_/weed_free_hay.html) As such, native, weed free hays should be sought. A list of certified weed-free hay (and potentially native hay) producers is usually available on the web for each state through Departments of Agriculture. Regardless, weed content of any material must be very carefully monitored and controlled by inspection and certification as written and/or required in project specifications. Straw is often used for mulch, but the almost inevitable inclusion of grain seed in straw materials makes the use of straw highly inadvisable because of the competition that results from germinating grain plants. HYDROMULCH Hydromulch is essentially wood and/or paper fibers that are mixed into a water slurry and sprayed onto the ground surface to apply a mulch coating of varying thickness. Hydromulch should be applied separately following seed application. The hydromulch mix is often colored green to assist operators in applying an even cover during spraying. The green color usually fades to tan or gray within a few weeks. An organic tackifier is typically added to the slurry to enhance the durability of the applied mulch cover. Although typically applied at a rate of approximately 1,500 pounds per acre (1680 kg/ha), it is more effective at a rate of 3,000 pounds per acre (3360 kg/ha) with a guar gum tackifier. Although hydromulches are effective, they are substantially more expensive than other mulch types. BONDED FIBER MATRIX Bonded fiber matrix is a spray-on mat consisting of a continuous layer of elongated fiber strands held together by a water-resistant bonding agent which creates a very durable and ground-fitting cover. Bonded fiber matrix is especially useful where steep and very rocky surface conditions would make the use of erosion control mats ineffective. It is important to assure a trained technician applies the bonded fiber matrix as a continuous cover is needed to create the integrated shell, but if the material is applied too thickly it can prevent penetration of seedling shoots. EROSION CONTROL MATS There are a large variety of mats which can perform the function of mulch. Mats composed of aspen shavings attached to or sandwiched between one or two plastic nets have long been used with good success. There are also many mats comprised of different combinations of coconut fiber, straw and other materials that double as both mulch and erosion control. Erosion control mats are generally expensive and are most cost-effective when used on areas where erosion potential is high and the site surface is relatively smooth. Use of erosion control mats on rough ground is less effective because the mulch to ground contact is poor. Where the surface is very rocky, material ends up suspended above most of the surface stretched between protruding rocks. Some mats have unsightly and non-environmentally friendly plastic netting that does not biodegrade and can be stripped off by wind and/or causes issues with wildlife getting caught in them. Biodegradable netting (not to be confused with “photodegradable” plastic netting) is available from some manufacturers at a slightly higher cost. SAND OR ROCKS In harsh sites, relatively thick layers or sand or small rocks can also serve to conserve moisture as well as to hinder weed establishment. PLANTING Planting can complement seeding efforts and increase the overall success of a restoration project and should not be overlooked because of the initial higher cost. Planting may be most desirable when desired species are difficult to establish from seed, unavailable, or the only seed sources available have low germination rates. Planting can be more effective when the revegetation site has abnormally stressful environmental conditions, such as extremely low nutrients, alkalinity, salinity, erosion potential or a short growing season, provided proper irrigation is supplied, when the existing plant community will present severe competition during seedling establishment, or a need for more rapid plant establishment exists than can be met through seeding. If planting is limited by budget constraints, fewer plants may be used by creating islands of more mature plants to produce a more diverse and natural looking landscape as well as a central, established stands of native plants that can reproduce and spread. Care must be taken to protect plants as needed to assure over-browsing does not occur. At many sites, protective tubes or cages should be installed on trees and shrubs to protect them from wildlife predation until they are well established. Plant Materials Bareroot and containerized plants are common types of transplant stock usually purchased from a nursery. If you are purchasing plant materials from a commercial supplier, the decision to use bareroot or containerized stock should be based on species characteristics (Shallow vs deep rooted species), site conditions, scheduling and cost. Larger containers are more expensive to purchase, transport, store, handle and transplant. However, post-transplant growth has been shown to increase with container sizes, provided proper irrigation is supplied. In general, larger stock performs better under more adverse site conditions (Landis and Simonich 1984). Ideal ages of containerized stock herbaceous plants 1 to 3 years woody plants/shrubs 3 to 5 years trees 5 to 10 years Ordering and Delivery Order bareroot or containerized seedlings from 1 to 4 years in advance of the planting date. In general, containerized seedlings should be ordered from between 1 to 2 years prior to planting; bareroot seedlings should be ordered from 2 to 4 years prior to planting (Townsend et al. 1993, Shaw 1984). Advanced planning and ordering will ensure the availability of desired species and proper hardening of the stock. In addition to their standard stock, some nurseries offer custom growing and may be able to propagate materials that are collected from the project site. However, custom grown materials have a higher initial purchase cost and may require additional production time as well as an initial contract and down payment (Townsend et al. 1993). Upon receipt, all containerized stock should be examined to ensure It has a root system adequate to hold the ball together but is not root bound. It is adequately hardened. The plant nursery should be held responsible for providing plants that are adequately hardened and should be willing to certify that greater than 80% of the plants are hardened to withstand temperatures of 20F for 8 hours. Visual observations of lack of hardening will include brown necrotic stem tissue, copious leaf drop, and brown, necrotic buds. Additionally, lush tender growth should be minimal and woody stem development should be occurring at the crown. Rooting must ample enough for the plug to retain its shape and potting mix after pulling. The photos below help illustrate acceptable vs unacceptable traits for planting (photos courtesy of NRCS Technical Notes State of Washington, February 2005) Hardy sage plant, note the high degree of root development when compared to the amount of top growth. Leaves are compact and turgid. Sage plant lacking cold-hardiness characteristics. Note elongated, supple light-green leaves. Rooting is ample, but woody tissue development is nonexistent and the amount of lush top-growth is disproportionately high. Other potential problems can be indicated by: Elongated buds. Leaf emergence. Elongation of white root buds. Root molds (not to be confused with mycorrhizae). Seedlings should also be examined for their moisture level, which can be determined by observing the root condition, the twig, needle or root flexibility, and the overall appearance (Townsend et al. 1993). Any damage or deficiencies in the stock should be immediately reported to the nursery and/or contractor from whom it was received. If deficiencies occur in the number of plants shipped or the packaging or transportation procedures, the nursery should be able to correct the problem. If serious problems are noted for the received materials, they should not be planted and should be classified as cull stock. Again, in such case, contact your nursery and/or contractor to correct the problem before payment is made (Townsend et al. 1993). SALVAGE An alternative to purchasing plants is to collect them in the wild (refered to herein as “salvage”). Salvaging native plants is ideal because 1) locally-adapted plants are “recycled,” 2) success of transplant survival can be high when the native soil accompanies the salvaged plant, and 3) salvaged plants can be more cost effective than purchased plants, especially for projects with limited budgets but plenty of labor (paid or volunteer). Salvaged plants can either be directly transplanted onto another site or potted and tended until the next planting season. Acquisition of Salvaged Material Salvaging from areas that have noxious weeds or undesirable plants should be avoided. Native plants or sod on sites slated for development can be salvaged. This method requires some advanced knowledge of construction areas and planning for the destination of salvaged plants. Small numbers of plants may be transplanted from sites ecologically similar to the revegetation site. On the other hand, if the revegetation project occurs in the same place as is being disturbed, plants can be salvaged from the site itself. All salvaged plants and associated soil should be weeded to prevent contamination from the donor site. WHAT TO SALVAGE General guidelines as to potentially salvageable material include:. Use plants that reproduce through vegetative sprouting (root shoots) Native plants that grow in disturbed areas are likely particularly suited for transplanting Do NOT try to salvage plants with taproots and/or extensive root systems Avoid diseased or weak plants WHEN TO SALVAGE The ideal time to salvage plants is during dormancy (October through April). However, plants can be salvaged at other times of the year if one-third to two-thirds of the plant is cut back and a good rootball is saved and kept moist. It is best to dig up plants in the morning and transplanting on another site or potting the plants in the afternoon. If plants are dug up when dormant, they should be “heeled in” in a pile of mulch or soil, and kept moist until they are transplanted. Dormant plants that are potted will need to be watered periodically through the dormant season. A soaking once every three weeks should be sufficient in cool weather. Increase waterings during warm spells. Transport plants in plastic grocery bags or moist burlap bags to conserve water. Heavy machinery can be used to move large areas of sod or clumps of shrubs. Cuttings, sodding, and plant salvage are less commonly used for upland species reintroduction, as containerized or bareroot stock is more suitable for these sites. Appendix X. Drill Calibration Summary (from Nancy Shaw, USFS Research Station)