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BI302 – Evolution Fall 2016 Instructor: Office: Telephone: E-mail: Office Hours: Dr. S. M. Ramsay N3022C 884-0710 x2406 [email protected] M 14:00 – 16:00 R 12:00 – 14:00 Lectures: Tutorials: 3 hours 1.5 hour MWF 8:30 – 9:20, BA202 T1: T 4:00 – 5:20, N1042 T2: T 5:30 – 6:50, N1042 T3: R 5:30 – 6:50, N1042 T4: R 4:00 – 5:20, N1042 T.A.: Ms. Kirsten Reid [email protected] T.A.: Ms. Megan Warkentin [email protected] TA: Megan Warkentin TA: Megan Warkentin TA: Kirsten Reid TA: Kirsten Reid IA: Claire Watterson IA: Claire Watterson IA: Dane Goh IA: Dane Goh Prerequisites: BI226 – Genetics Course Description From the Course Calendar: “A comprehensive and integrative course on evolution by natural selection as the underlying principle of modern biology. Topics include the mechanisms of selection; the concepts of adaptation, fitness and species; the evolution of sex; co-evolution; and the origin of life.” From The Origin of Species: “It is interesting to contemplate an entangled bank, clothed with plants of many kinds, with birds singing on the bushes, with various insects flitting about, and with worms crawling through the damp earth, and to reflect that these elaborately constructed forms, so different from each other and dependent on each other in so complex a manner have all been produced by laws acting around us….and, that whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been, and are being, evolved.” In this course you will learn about the processes (Darwin’s “laws acting around us”) involved in the evolution of populations of organisms. In addition to learning concepts you will also develop your critical thinking skills, and your verbal and quantitative reasoning; you will work at thinking like a scientist. Textbooks Required Zimmer C and Emlen DJ, 2015. Evolution: Making Sense of Life, Second Edition. Roberts and Company. Perkins AEH. 2015. Study Guide to Accompany Evolution: Making Sense of Life, Second Edition. Roberts and Company. Pechenik JA. 2015. A Short Guide to Writing about Biology, 9th Edition. Pearson-Longman Publishers Recommended Supplemental Texts Darwin, C. 1859. The Origin of Species, Second Edition. Available online at http://darwin-online.org.uk Additional readings As assigned. Evaluation Thought problems (best 3 out of 4) Quizzes (best 3 of 4, during tutorial) Participation Midterm Exam (Saturday 29 Oct, 9:00 am, TA) Final Exam (final exam period in December) Percentage Letter Grade 90-100 A+ 85-89 A 80-84 A- 77-79 B+ 73-76 B 70-72 B- 67-69 C+ 63-66 C 60-62 C- 57-59 D+ 53-56 D 50-52 D- 0-49 F 20% 15% 10% 20% 35% Grade Definition Excellent Good Adequate Marginal Inadequate Strong evidence of original thinking; good organization; capacity to analyze and synthesize; superior grasp of subject matter with sound critical evaluations; evidence of extensive knowledge base. Evidence of grasp of subject matter, some evidence of critical capacity and analytic ability; reasonable understanding of relevant issues; evidence of familiarity with literature. Student who is profiting from his/her university experience; understanding of the subject matter; ability to develop solutions to simple problems in the material. Some evidence of familiarity with subject matter and some evidence that critical and analytic skills have been developed. Little evidence of even superficial understanding of subject matter; weakness in critical and analytic skills; with limited or irrelevant use of literature. Late Assignments and Missed Quizzes Thought problems will be accepted up to a maximum of 72 hours after the due date. A late penalty of 10% will apply for each 24-hr period the assignment is late. There will be no makeup for missed quizzes. Information Technology and Electronic Devices You and I will be making extensive use of the resources on MyLearningSpace. Important dates for the course, including assignment deadlines, are listed on the calendar. I will post documents and links on the content page. The discussion forums are available to ask and answer questions, and share information with everyone in the class. You will submit all of your assignments through the dropbox. Your grades will be regularly updated and visible so you can track your performance. If you can think of other ways we can make good use of MyLS, I am always happy to hear about them. The use of information technology, and electronic devices in the classroom are governed by University policies outlined in the academic calendar: http://www.wlu.ca/calendars/section.php? cal=1&s=568&ss=2429&y=57. I expect you to use technology and devices for productive purposes. Any use that is disruptive to the academic environment of the classroom is prohibited. I will ask offenders to leave the class, and I will report offenses to the Dean of Science. Lectures The lectures in this course (3 hours per week) are intended to help you work through the material in the textbook. Evolution (both the topic and this course) is one of the most conceptual subdisciplines in biology; this is not a course where you can expect to be successful by cramming. As such, it is crucial that you keep up with the readings from the textbook. In addition, as we move through the material, there may be occasions where sections of the text are assigned for you to read through on your own (e.g. during the first week of class you should read through Chapter 5 and test yourself with the questions at the end of the chapter and in the study guide as a refresher of the material you learned in BI226). Each of the chapters builds heavily on the previous ones, so retention of concepts is essential and falling behind early in the course will have exaggerated effects later on. In addition to the readings, you should work through the questions at the end of each chapter, and the exercises in the accompanying study guide. Tentative Lecture Schedule Week 1 (5 – 9 Sept.) Monday Labour Day No Class Wednesday Orientation No Class 2 (12 – 16 Sept.) Ch. 1 & Ch. 2 (Introduction to Evolutionary Biology) Ch. 6 (Drift and Selection) Ch. 6 & Ch. 7 (Population & Quantitative Genetics) Ch. 7 (Phenotypic Evolution) Ch. 4 (Phylogenetics & Tree Thinking) Ch. 6 (Drift & Selection) Ch. 7 (Quantitative Genetics) Friday Ch. 1 & 2 (Introduction to Evolutionary Biology) Ch. 4 (Phylogenetics & Tree Thinking) Ch. 6 (Population Genetics) Ch. 7 (Phenotypic Evolution) Ch. 8 (Natural Selection in Action) No Class Ch. 8 (Natural Selection in Action) No Class Ch. 9 (Coalescence & Molecular Phylogenetics) Ch. 11 (Sex & Sexual Selection) Ch. 11 (Sex & Sexual Selection) Ch. 12 (Life History Evolution) Ch. 16 (Behavioural Evolution) Ch. 13 (Speciation) Ch. 15 (Coevolution) Ch. 9 & Ch. 10 (Genomic Evolution & Adaptation) Ch. 11 (Sex & Sexual Selection) Ch. 12 (Life History Evolution) Ch. 16 (Behavioural Evolution) Ch. 13 (Speciation) Ch. 15 (Coevolution) Ch. 14 (Macroevolution) Conclusion Study Day No Class 3 (19 – 23 Sept.) 4 (26 – 30 Sept.) 5 (3 – 7 Oct.) Fall Reading Week (10 – 14 Oct.) 6 (17 – 21 Oct.) 7 (24 – 28 Oct.) 8 (31 Oct. – 4 Nov.) 9 (7 – 11 Nov.) 10 (14 – 18 Nov.) 11 (21 – 25 Nov.) 12 (28 Nov. – 2 Dec.) Thanksgiving No Class Ch. 9 (Coalescence & Molecular Phylogenetics) Ch. 10 (Adaptation) Ch. 11 (Sex & Sexual Selection) Ch. 12 (Life History Evolution) Ch. 16 (Behavioural Evolution) Ch. 13 (Speciation) Ch. 15 (Coevolution) 13 (5 – 9 Dec.) Ch. 14 (Macroevolution) Tutorials You will have one-and-a-half-hour tutorials weekly on Tuesday or Thursday afternoons. These sessions are intended to supplement the lecture material and will involve two different types of exercises: thought problems, and problem solving. Thought problems will involve discussion of selected readings with questions that will be assigned one week before the tutorial. There are four of these sessions scheduled; for each of these you will write a short (2 page) answer to the thought problem. The thought problems will help you to enhance your skills of reading and analysing scientific papers, and making clear, logical arguments in support of a position. The problem solving tutorials are designed to help you work through some of the quantitative and analytical aspects of evolutionary biology. There are six of these scheduled; the goal for each is to develop your quantitative reasoning skills, and to give you practice with problems specific to evolutionary biology. Four of the problem solving sessions will include quizzes that will test the skills you have developed over the previous few weeks and will give you some additional preparation for the midterm and the final exam. You should have begun working on some of the end-of-chapter and study-guide problems prior to the tutorial and come to class with questions. Tutorial Schedule Week 2 (13/15 Sept.) 3 (20/22 Sept.) 4 (27/29 Sept.) 5 (4/6 Oct.) 6 (18/20 Oct.) 7 (25/27 Oct.) 8 (1/3 Nov.) 9 (8/10 Nov.) 10 (15/17 Nov.) 11 (22/24 Nov.) 12 (29 Nov./1 Dec.) Exercise Intro to your TA Problem Solving, Quiz Tentative Topic Effective Writing for Thought Problems; Chapter 1 & 2 Review Conceptions and Misconceptions of Evolution Phylogenetic Trees & Population Genetics Problem Solving Quantitative Genetics/Phenotypic Evolution Problem Solving, Quiz Natural Selection Thought Problem 2 Neutral vs Adaptive Evolution Problem Solving Adaptation and Complex Traits Thought Problem 3 Sexual Selection Problem Solving, Quiz Thought Problem 4 Life History Variation/Components of Fitness Behavioural Evolution Problem Solving, Quiz Speciation, General Review Thought Problem 1 Participation Everyone is expected to participate in the tutorial activities. Most of the work will be group work, and the TAs will be monitoring your involvement as you do your work. When it comes time to speak up, everyone is expected to take a turn, and again the TAs will be making note of your contributions. BI302 Evolution Learning Objectives On completion of this course, students will be able to 1. demonstrate an in-depth understanding of how the scientific method, and the central organizing paradigms, concepts, and processes, contribute to knowledge about evolutionary biology (Biology Program-level Learning Outcome 1); 2. articulate an understanding of how evolutionary biology integrates with concepts and procedures from across the natural sciences, to contribute to the development of new understanding of life processes (Biology Program-level Learning Outcome 2); 3. select and apply concepts, theories, and established methodologies currently used in evolutionary biology for the identification, investigation, and analysis of complex problems (Biology Program-level Learning Outcome 3); 4. propose and test concepts, hypotheses and predictions about evolution using critical thinking and analytical skills at all steps in the process (Biology Program-level Learning Outcome 6); 5. communicate information about evolutionary biology, from a variety of sources, in a variety of forms, including informal and formal reports, to a range of audiences (Biology Program-level Learning Outcome 7); 6. evaluate current evolution research in the context of the needs of society in areas such as conservation, environmental quality and health, and the ethical issues that may be involved (Biology Program-level Learning Outcome 9) IMPORTANT NOTES Adding and Dropping Important Dates for Course Adding/Dropping and Tuition charges for the Fall 2016 Term, please see this link: https://legacy.wlu.ca/calendars/dates.php?cal=1&t=284&y=69 Important Dates for Course Adding/Dropping and Tuition charges for the Winter 2017 Term, please see this link: https://legacy.wlu.ca/calendars/dates.php?cal=1&t=285&y=69 Examination Deferrals The Academic Date section of the Calendar (Printed and Web Site Versions) clearly states the examination date period for each semester. Students must note that they are required to reserve this time in their personal calendars for the examinations. The Fall examination period is: December 10‐21, 2016. The Winter examination period is: April 6‐24, 2017. Students who are considering registering to write MCAT, LSAT or GMAT or a similar examination, should select a time for those examinations that occurs outside the University examination period. For additional information that describes the special circumstances for examination deferment, please check the following web page: https://students.wlu.ca/academics/exams/deferred‐exams.html – Faculty of Science: Request for Deferred Final Examination. Student Awareness of Accessible Learning Centre Students with disabilities or special needs are advised to contact Laurier’s Accessible Learning Centre for information regarding its services and resources, ext. 3086. Students are encouraged to review the Calendar for information regarding all services available on campus. https://alc.wlu.ca/. Academic and Research Misconduct Academic misconduct is an act by a student, or by students working on a team project, which may result in a false evaluation of the student(s), or which represents a deliberate attempt to unfairly gain an academic advantage. For detailed information on Academic Misconduct please refer to: https://students.wlu.ca/academics/academic‐integrity/index.html. Plagiarism Detection Software Wilfrid Laurier University uses software that can check for plagiarism. Students may be required to submit their written work in electronic form and have it checked for plagiarism. Learning Services There is a range of academic learning support services offered at Laurier designed for all students who want to improve their academic achievement in the classroom. These services include the following specific areas: * Central Academic Advising Office * Mathematics Assistance Centre * Study Skills and Supplemental Instruction Centre * Writing Centre Please access the following web page for detailed information: https://students.wlu.ca/academics/support‐and‐advising/index.html. Laurier Email Account Our official means of communication is with your Laurier email account. Students are expected to regularly check their Laurier email account for important notices from the university community. Students are also expected to send emails to official members of the university community from their Laurier email account in order to ensure delivery. Emails sent from non‐Laurier accounts, such as hotmail, may be identified as spam and not delivered. Your co‐operation is appreciated.