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Darwin and Speciation Notes Meet Charles Darwin • 1809-1882 • Published On the Origin of Species in 1859 • Fun fact! Darwin dropped out of medical school after seeing his first cadaver Darwin’s Findings • Darwin observed many species during his voyage on the HMS Beagle • In the Galapagos, Darwin observed the variations of animals and plants from island to island. • This eventually led to his theory of evolution by natural selection Building on Earlier Science • Darwin built his theory off the work of JeanBaptiste Lamarck • Lamarck wrote that living things change over time • Lamarck thought that organisms passed on their most-used traits to their offspring – Is this true? No! What is a Species? A species is a group of similar individuals capable of interbreeding to produce fertile offspring. How Many Species Are There? We don’t know Estimates range from 4 to 100 million ~2 million species have been described How Do Species Arise? Speciation- evolutionary process by which reproductively isolated biological populations evolve to become distinct species. Let’s break that down, shall we? First things first: Evolution = ? Change over time! How Do Species Arise? The key to speciation is reproductive isolation of populations. Geographic isolation is most common Process of speciation: Parent species I. Barrier 2 separate species II. Diverge and Mutate III. No Longer Able to interbreed Two Types of Speciation 1. Allopatric speciation: geographic isolation creates a reproductive barrier (an extrinsic mechanism). Allopatric Speciation Example Harris’ antelope squirrel White-tailed antelope squirrel Two species of ground squirrel are thought to have descended from a common ancestral population that was separated by formation of the Grand Canyon. Two Types of Speciation Sympatric speciation: reproductive barrier is created by something other than geographic isolation (intrinsic mechanism). Sympatric Speciation Example Ancestors of apple maggot flies laid their eggs on hawthorns, a plant species native to North America. When immigrants came from Europe, they brought apple seeds. Apple maggot flies now had a choice of where to lay their eggs—hawthorns or apples. Flies hatched on apples only mate with applehatched flies, while flies hatched in hawthorns only mate with hawthorn-hatched flies. How Fast Does Speciation Occur? It depends! Speciation relies on many factors Examples: A slow rate of speciation evidenced by a living horseshoe crab (13 extant species) and a 300 million year-old fossil species. In contrast, Galapagos finches have diversified into 13 species within the last 100,000 years. Does Evolution Create the Perfect Organism? No, only better organisms as evolution is constrained by history and buffeted by random events. What do we mean by “better?” Better for survival and reproduction