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WORKSHOP REPORT 3 Management of moorland edge for priority species Report of a Workshop held at ADAS Pwllpeiran 17 June 2003 1 Contents BACKGROUND 4 MOORLAND EDGE HABITATS: DEVELOPMENT AND DISTRIBUTION - TIM MILSOM (CENTRAL SCIENCE LABORATORY) 4 Description of moorland edge habitats 4 Development of moorland edge habitat 5 Distribution and composition of the moorland edge 6 Moorland edge and conservation of species and habitats Vegetation Invertebrates Reptiles and Amphibians Birds 8 8 8 9 9 Summary 9 MOORLAND EDGE MANAGEMENT – CLARE BURROWS (COUNTRYSIDE COUNCIL FOR WALES) 10 Degraded Fridd zone 10 Management of specific habitats Bracken Woodland Arable land 11 11 11 12 Summary 12 MANAGEMENT OF MOORLAND EDGE FOR BIRDS - SIAN WHITEHEAD (COUNTRYSIDE COUNCIL FOR WALES) 13 Introduction 13 Community and species associations with Moorland Edge Habitat associations identified by this study Species associations based on other studies Questions to be addressed 13 13 13 14 Fridd management 14 Management of moorland adjacent to forest edge Experience from the RSPB black grouse recovery project 14 14 Summary 15 2 FIELD TRIP 16 Brignant biodiversity paddocks 16 Oak woodlands 16 Parc Vicarage wetland 16 Hafod Grasslands 17 Ty Coch bracken and scrub 17 SUMMARY 17 ACKNOWLEDGEMENTS 18 AUTHOR CONTACTS 18 REFERENCES 18 3 Background This workshop was held at ADAS Pwllpeiran in Cwmystwyth, and focused on the conservation management of moorland edge. The workshop was prepared for land managers and project officers with responsibility for these habitats and included formal talks, informal discussions and a field visit to moorland edge habitats. The following topics were discussed at the workshop: The definition of moorland edge The extent of moorland edge Examples of moorland edge How moorland edge habitats have developed Management of moorland edge for plants, birds and other species Moorland edge habitats: development and distribution - Tim Milsom (Central Science Laboratory) Description of moorland edge habitats Moorland edge is described as the zone of transition between improved agricultural land and moorland. It has characteristics of both intensively managed land and semi-natural moorland. Pockets of moorland may be found within the moorland edge zone, whilst conversely, pockets of moorland edge habitat can exist within moorland. Typical habitat types are listed in Table 1. Classification of parcels of land into the moorland edge is not always obvious, however, and may depend upon the size of the parcels and their adjacent habitats. Some features, such as plantation forests or reservoirs, are large enough to be considered features in their own right. Also there is no clear definition of moorland edge based on vegetation type alone; thus large areas of unmanaged grass dominated by species such as purple moor grass (Molinia caerulea) or mat grass (Nardus stricta), may be considered to be moorland rather than moorland edge. One of the most important characteristics of moorland edge is that it is a mosaic of small habitat patches. The edge also tends to be more diverse than the surrounding agricultural land, grassland or heather moorland (Table 1). 4 Table 1: Habitat composition (mean % cover) of moorland edge and adjacent habitats in 12 uplands of England and Wales Habitat Wood – deciduous Wood - mixed broad leaved Conifers – mature Conifers -felled/new plantation Scrub /gorse Bracken Dwarf shrub heath - dense ericaceous Dwarf shrub heath - open Agricultural Moorland Grassland Heather edge moorland 8 9 1 1 2 3 . . 2 16 5 2 . 1 . . . . . . . 7 4 2 . . 1 23 . 1 3 24 Grass -improved (intensive) Grass -improved (hay/silage) Grass -neutral (rough) Grass -neutral (grazed) Grass – calcareous (rough) Grass – calcareous (grazed) Grass – acid Grass – acid (rough) Grass – acid with Juncus Grass – acid with Nardus/Festuca/Molinia 51 9 1 2 3 3 1 . . . 2 . 2 25 9 1 12 1 1 5 . . . 9 2 . 44 1 . 26 . . . 5 1 . 16 . . 8 Arable Set-aside Bare ground Fen / marsh Bog 16 . . . . . 2 . . . . . 1 . 12 1 . Development of moorland edge habitat1 Moorland edge communities are the product of alternate phases of agricultural improvement and abandonment which have been occurring since the 11th century, at least. The changes in landuse along the moorland edge often give rise to a mosaic of habitats, at various stages of improvement, e.g. reseeded pastures or small blocks of alien conifers, or abandonment e.g. bracken beds, or scrub.. The management practices which have had the 1 Much of the survey work has been carried out by Parry et al. of Birmingham Univerity, and by Silsoe College. 5 greatest effect on the dynamics of the moorland edge have been, and continue to be, grazing (including density, timing and species), agricultural improvement, agricultural reversion and afforestation. In the 19th and 20th centuries particularly, areas of rough grassland in the lower parts of the moorland edge were agriculturally improved by draining, liming, fertilising, and then enclosing them. Reversion took place as the drains became blocked and inputs reduced; rough grass, bracken (Pteridium aquilinum) and scrub then invaded. This reversion, which was particularly prominent in the Yorkshire Dales and Dartmoor, lasted until the 1970s and 80s. At this time agricultural improvement became widespread in some areas, namely, central Wales and the Yorkshire Dales, Lake District and Snowdonia National Parks though the ‘improvements’ were less evident in other areas. The situation in the 1990s and beyond is not yet clear, but it should become so as data from Environmentally Sensitive Area (ESA) monitoring programs become available. At the upper edge of the transition zone, moorland may be lost to moorland edge communities through excessive burning or over-grazing by livestock. For example, burnt heather moor might be invaded by bracken, whereas localised overgrazing or winter grazing by livestock may result in loss of dwarf shrub and an increase in rough grasses. The tendency in recent decades to graze sheep only rather than sheep and cattle may also have contributed to the loss of species and diversity. The planting of non-native coniferous forests has been evident particularly in Northumberland, North York Moors, Snowdonia and the Brecon Beacons. It has tended to obliterate the moorland edge transition zone as few species survive in dense forests except along the forest rides. Distribution and composition of the moorland edge Using satellite data from the Land Cover Map 2000, estimates of moorland edge in 12 upland regions of England and Wales have been made (Fig. 1). The total area of moorland edge in the 12 regions is thought to be 680,175ha, ranging from 5,812ha in the Shropshire Hill ESA, to 148,425ha in the North Pennine Dales. The proportion of hill edge to moorland ranged from an estimated 31% in the North York Moors to 127% in the Shropshire Hills and 203% in the Lake District. This latter estimate is considerably higher than for the other regions and there is some concern about its accuracy. 6 Region Area of hill-edge as % of moorland area 0 50 100 150 200 250 North Pennines Snowdonia South Pennines Lake District Mid-Wales Brecon Beacons Northumberland Dartmoor Bowland North York Moors Exmoor Shropshire Hills 0 20 40 60 80 100 120 Area of hill-edge (ha * 1000) Area 140 160 % of moor Fig1: Estimates of moorland edge in 12 upland regions of England and Wales Figure 2 shows that there is significant variation in habitat composition across the 12 upland areas of England and Wales. Natural and semi-natural grassland communities dominate the moorland edge. Most of these are acidic grasslands but calcareous grasslands are important in the North Pennines, Bowland and the Shropshire Hills, and neutral grasslands in the North Pennines, Bowland and Northumberland. Conifers are a dominant feature in Northumberland and the North York Moors, whilst deciduous woodland cover is relatively high in Exmoor, Dartmoor and the North York Moors. Scrub occurs in small, scattered areas and appears to be under threat. It includes the BAP (Biodiversity Action Plan) priority species, juniper (Juniper communis subsp. communis), stands of which can be found in the North Pennines. Continuous stands of bracken (which are detectible from satellite images) comprise less than 5% of the moorland edge communities; they appear to be particularly extensive in the Lake District and has increased in recent years. Dwarf shrub heath, improved grassland and arable land are generally seen as minor components of the moorland edge habitat in all areas. 7 Region Northumberland Lake District North Pennines North York Moors Bowland South Pennines Shropshire Hills Snowdonia Mid-Wales Brecon Beacons Exmoor Dartmoor Deciduous wood Conifers Scrub & bracken Dwarf shrub heath Nat. & seminat. grass Improved grass Arable 0 20 40 60 80 Cumulative % cover 100 Other Fig 2: Regional variation in habitat composition of moorland edge Moorland edge and conservation of species and habitats The amount of information that is available on plant and animal species and their requirements, and consequently their management, varies between species groups. Generally, there is not enough information to form detailed management plans except for a few targeted species such as black grouse (Tetrao tetrix). Many of the species found on moorland edge can also be found in other areas, and others do not use the moorland edge exclusively for all of their activities. Below is an overview of conservation issues within different species groups. Vegetation Many of the vegetation communities of importance have been described in previous sections of this report. Potential moorland edge communities for which there is a BAP include upland heathland, upland calcareous grassland, upland mixed ash woods, native pine woodlands, wet woodland, limestone pavements and blanket bog. Maintaining the mosaic is probably more beneficial for biodiversity than protecting individual communities, except perhaps where BAP priority species are an issue. Juniper (Juniperis communis subsp. communis) for instance, is under threat primarily because of high stocking rates. Juniper should be grazed lightly in summer only. Short swards with gaps are required for seedling establishment and this has been most successful when grazing pressures have varied over time. Invertebrates Upland invertebrates are more diverse than upland plants and mammals. They benefit from the mosaic of habitats as well as from variations in plant structure and topography. 8 Wet flushes may also be beneficial. Management that is detrimental to invertebrates includes agricultural improvement, heavy grazing and afforestation as these practices reduce diversity and structure. BAP species include High brown fritillary (Argynnis adippe) and Pearl bordered fritillary (Boloria euphrosyne). Management to promote these is described later in this report. Reptiles and Amphibians Little is known of the requirements of these species in the uplands though it is thought that the common frog, the common toad and palmate newt require wet habitats. Other species include the lizard and adder and none of them are found exclusively on moorland edge. Birds Two species of birds, Black Grouse (Tetrao tetrix) and Red Kite (Milvus milvus) (Welsh population only), are specialists of the moorland edge, whilst several other species, which nest on moorland or cliffs, use it as a feeding habitat. This latter group includes: Golden plover (Pluvialis apricaria) Curlew (Numenius arquata) Ring ouzel (Turdus torquatus) Twite (Carduelis flavirostris) Merlin (Falco columbarius) Golden eagle (Aquila chrysaetos) Moorland edge may also act as a refuge for several lowland bird species that are in retreat, such as: Lapwing (Vanellus vanellus) Snipe (Gallinago gallinago) Whinchat (Saxicola rubetra) Most moorland edge species have declined in England and Wales, red kite and twite are exceptions. The causes of this are not fully understood, but it is believed that agricultural improvement has a negative affect on many bird populations, particularly ground-nesting species. Also, whilst young coniferous plantations might be exploited by black grouse, short – eared owl (Asio flammeus), hen harrier (Circus cyaneus) and whinchat, they will be deserted as the canopy becomes more dense. This subject is treated in more detail later in the report. Summary The moorland edge landscape has been shaped by a cycle of agricultural improvement and reversion. This has created the typical mosaic of communities that is important for the promotion of biodiversity, though many species associations with moorland edge are poorly understood. Agricultural improvement, which is thought to be detrimental to 9 moorland edge biodiversity, was widespread in the 1970s and 80s whilst data for more recent years is not yet available. It is hoped that the advent of ESA, Countryside Stewardship and Tir Gofal Schemes will have reversed the trend of the 1980s. There is, perhaps a need for prescriptions within these schemes that promote and maintain the mosaic of habitats found within the moorland edge zone. Moorland edge management – Clare Burrows (Countryside Council for Wales) Moorland edge can be a very variable land type and in Wales, is commonly known as Fridd land. The following is a list of all habitats larger than 1km2, found on Welsh moorland edge. The habitats are listed in order of frequency, but the list is not exhaustive. Improved grassland Acid grassland Dense bracken Conifer plantation Broadleaved woodland Acid dry heath Juncus dominated wet grassland Semi-improved wet grassland Molinia dominated wet grassland Acid flush Wet heath Scrub Neutral grassland Moorland edge typically consists of a higher diversity of habitat than the moorland per se, and it is believed that the moorland edge supports a diversity of species, though research is still in its infancy. The majority of the species that utilise moorland edge are not specialist to one habitat but exploit the diversity of habitats present. For example, the dark green fritillary butterfly (Argynnis aglaja), requires a mosaic of two-thirds wet grass to one-third bracken with adjacent scrub. This combination is now unlikely to be found in the lowlands and consequently moorland edge has become an important refuge for this species. Degraded Fridd zone The moorland edge has often degraded at the lower edge where land has been agriculturally improved (as described previously in this report). In these cases there is scope for landscape re-creation, but before any re-creation plan is embarked on, existing species should be considered. Short unfertilised swards, which are apparently species poor, may be home to wax cap fungi (Hygrophoraceae) for instance. If more than ten species of this fungi are present at a site it would be considered to be a site of national importance and clearly, such habitats should not be interfered with. 10 Management of specific habitats Bracken High brown fritillary and pearl-bordered fritillary are two BAP priority species that can be found in bracken. The high brown fritillary has declined dramatically in England and Wales since the 1950s, and is now found on about 50 sites. The butterfly used to be found in coppice woodland, much of which has now been lost to bracken communities, which do consequently have a similar flora to the coppice woodland. The species is now found on bracken slopes on moorland edge land in the west of the UK. It has declined partly because of a lack of management of bracken, for example a decline in the practice of cutting bracken for livestock bedding. Adult high brown fritillaries are on the wing from July to August, and require nectar sources such as thistles and brambles to be present close to their breeding sites. The eggs are laid within a few centimetres from the larval food plant. Since eggs overwinter, they require a substrate that is unlikely to rot, and bracken is ideal for this. The eggs hatch in spring and require warmth, which is provided by the litter, which is effective in trapping warm air. Larvae require an abundant food supply of violets, particularly the common dog violet (Viola riviana) which can be found in bracken litter. Habitat management for the high brown fritillary may include grazing with cattle or horses in winter and spring at a rate low enough for the litter layer to develop, but high enough to prevent bracken becoming dense enough to shade out the violets and to prevent grasses growing. Some light summer grazing may also be beneficial, though grazing by sheep is not suitable. The pearl-bordered fritillary has a similar ecology and habitat requirement to the high brown. Its range has also contracted out of coppice woodland onto bracken covered slopes, especially south facing ones. The adults fly from late March to May, and feed on bugle (Ajuga reptans), ground ivy (Glechoma hederacea) and blue bell (Hyacythoides non-scriptus) close to the breeding site. The larvae also feed on common dog violet. It has been suggested that these butterfly species exist in population clusters (metapopulations) i.e. a number of local populations connected by occasional dispersal. This feature may enable the butterflies to adapt readily to changes in habitat suitability, with populations becoming extinct in localised areas as habitat suitability wanes and colonising new areas of habitat as they become suitable. Creation of new areas of suitable habitat within range of a local fritillary population may help to enhance the persistence of these species. Woodland As mentioned previously there are a number of upland woodland habitats for which there is a Habitat Action Plan. Patches of woodland add to the biodiversity of the moorland edge not only in terms of plant species, but also because of the structure that contrasts with that of the grassier habitats and so provides habitats for a wider range of animals. 11 Agri-environment schemes encourage woodland to be fenced off. There are, however, four potential types of woodland management: Non-intervention in which grazing is excluded. Since herbivory is part of the natural woodland processes, exclusion can result in highly competitive species dominating the wood. High forest, in which mature trees are removed for timber Coppice. This is worth re-instating if there has been a recent history of coppicing i.e. in the last 20-30 years. Otherwise it is not recommended as long established species, such as lichens, can be damaged. Wood pasture, which is a traditional management regime. Under this regime, moderate grazing is permitted to prevent dominance by aggressive species. Establishing the correct stocking rate to maintain a diverse mixture of regenerating tree species is difficult. As a guide, it is suggested that within a 100 year cycle, the woodland should be grazed at 0.05 LSU (livestock units) ha-1 for 15 continuous years and the remaining years at 0.6 LSU ha-1. If management of such a wood has just been adopted, an assessment of the amount of tree regeneration already occurring should be made. If it is reasonably healthy then the 100-year cycle can be entered at the higher stocking rate. A particular disadvantage for this regime is that agri-environment scheme agreements are set for a maximum ten-year period only, which is too short a period for this regime. Arable land Arable cropping on the upland fringes used to be common, comprising up to one third of land in arable production. It has declined because conditions are less suitable than they are in the lowlands and profits were frequently uncertain. Arable cropping can add to the diversity of the moorland edge and has been encouraged under Tir Gofal and the Countryside Stewardship Scheme. In Tir Gofal the arable option has had a reasonable uptake thanks partly to the setting up of machinery rings, but also because a crop which fails to ripen can optionally be made into arable silage. Unsprayed arable crops on moorland edges are very valuable for nectar supplying plant species such as: shepherd’s-needle (Scandix pecten-veneris) corn marigold (Chrysanthemum segetum) cornflower (Centaurea cyanus) small flowered catchfly (Silene gallica) In addition, the overwintering stubble provides seed and is therefore a valuable extra food source for birds. Summary In conclusion, it appears that in order to enhance the biodiversity value of moorland edge habitats: 12 a diverse mix of habitats is important management should be planned at a landscape scale consideration should be given to what species already exist and their habitat requirements, before altering current management practices. Management of moorland edge for birds - Sian Whitehead (Countryside Council for Wales) Introduction Moorland edge is considered to be a transition community that is neither open moor nor semi-improved land. Moorland edge can consist of many diverse communities such as grass, bracken, gorse or hawthorn, on which a rich range of birds from the passerine, wader and raptor groups are wholly or partially dependant. Community and species associations with Moorland Edge Survey work carried out by the RSPB in the 1980s and recently analysed by the BTO2, has shown that there is a gradient in moorland edge community trends in Wales. In the north west, moorland edge communities tend to be dominated by grass or heather/bilberry (Calluna vulgaris/Vaccinium myrtillus) whilst those in the south east tend to be smaller and dominated by bracken. The survey also established some broad habitat and species associations with the moorland edge communities. Habitat associations identified by this study grass supports meadow pipit (Anthus pratensis), wheatear (Oenanthe oenanthe), skylark (Alauda arvensis) and ring ouzel bracken supports redstart (Phoenicurus phoenicurus), yellowhammer (Emberiza citrinella), tree pipit (Anthus trivialis) and whinchat (Saxicola rubetra) gorse supports stonechat (Saxicola torquata), dunnock (Prunella modularis) (hedge accentor) and linnet Species associations based on other studies Breeding waders such as curlew, golden plover and lapwing tend to favour more grassy areas for nesting and feeding. Wet flushes are also favoured for feeding in late summer. Groups of trees are avoided as these provide nesting and perching for corvid predators. Raptors such as red kite nest in woodland and hunt for carrion over the moorland edge. Ground nesting raptors such as hen harrier (Circus cyaneus), merlin and short eared owl (Asio flammeus) nest in the heather and hunt in grassier areas for voles and meadow pipits. Chough (Pyrrhocorax pyrrhocorax), which is a BAP high priority species, forages extensively on the acid grassland of Snowdonia. 2 The information presented here has been released by kind permission by RSPB and BTO on the understanding that it is to be used by POs for training purposes, but does not go beyond this report. 13 Questions to be addressed Are geographical differences a consequence of topography, climate, soil or land use? Can moorland edge be characterised? Can current vegetation be interpreted in terms of historic land use? What is the influence of mosaics on bird communities? What is the role of scrub and trees in bird-habitat relationships Fridd management There is a lack of knowledge concerning these edge communities (i.e. moorland edge not including coniferous forest) but they do appear to be important for birds. Until more information is available that would enable specific management prescriptions to be developed, they should be maintained by using appropriate grazing management, maintaining or creating wet areas, and preventing the use of fertilisers. Management of moorland adjacent to forest edge The edge of a coniferous plantation is a much simpler habitat than the Fridd communities. It is traditionally a hard, wind-firm forest stand abutting moorland and is potentially a valuable habitat for black grouse (Tetrao tetrix) and nightjar (Caprimulgus ruficollis), both of which are BAP priority species. For more information on the black grouse BAP, see www.gct.org.uk/research/blackgrouse Experience from the RSPB black grouse recovery project3 The black grouse recovery project started in 1999 as a response to a 50% decline of the Welsh population that has occurred since 1986. The BAP objective was to reduce the decline by half. Habitat management was undertaken in six key areas in mid and north Wales. The aim was to create and enhance favourable habitat at various spatial scales through a range of habitat prescriptions. Management for black grouse involves four habitat types: upland heath, bogs and flushes, bracken and forest. Upland heath Heather and bilberry provide food, cover and a habitat for black grouse to nest in. Cutting or burning maintains the heather at various stages; young shoots are nutritious whilst long heather is used for nesting. Drainage ditches should be blocked, possibly using brash from the areas of heather that have been cut. This will help to re-wet areas and prevent deaths of chicks which might otherwise fall in. Bogs and f lus hes Cotton grass (Eriophorum vaginatum) flowers are an important food source for the hens in spring. These wet areas are usually rich in insects that are eaten by hens and broods in summer. 3 The information presented here has been released by kind permission by RSPB on the understanding that it is to be used by POs for training purposes, but does not go beyond this report. 14 Brac k en Bracken provides cover but is of little dietary value, and control should be considered if it is an invasive threat. Fores t Forest coups (blocks of trees) with 15-40% canopy are preferred by black grouse. These provide sufficient understory for feeding whilst minimising the risk of predation since predators require cover. Forest thinning can be carried out at the forest edge in order to create a mosaic of partly covered and open areas. Trees should be felled such that the forest becomes gradually thinner as it approaches the moor edge, it will then blend into the moor rather than having the undesirable hard edge. Table 2 shows a suggested scheme for thinning within the forest and planting on the moor. Table 2: Idealised plan for thinning forest edge Open moor Tree cover (%) Density (trees/ha) Distance from open moor (m) Conifer plantation Band1 Band 2 Band 3 Band 4 10 20-50 60-80 100 440-1200 >1320 2200 41-85 86-100 50 broad 220 leaved 0-40 This management should only be carried out on young plantations since older stands are wind firm and felling within them could result in many other trees falling. The cut brashings can be used to block drainage ditches. On the moor, broad leaved trees can be planted at a rate of less than 50ha-1. This management may also benefit nightjar (Caprimulgus europeus). Summary When managing moorland edge it is recommended that mosaics of communities and gradients of successional habitats are maintained or created. This will optimise breeding and feeding areas for bird assemblages, whilst their loss will result in loss of the associated bird communities. 15 Field trip The aim of the field trip was to visit a selection of moorland edge habitats and discuss their management. Brignant biodiversity paddocks This is a species poor grassy area which was ‘improved’ in the late 1950s under the Cornhill improvement scheme which encouraged upland areas to be ploughed and reseeded with rye grass and white clover. It then continued to have annual dressings of fertiliser. In 1996 an experiment was set up to investigate ‘methods of maintaining and enhancing botanical diversity in upland ESA swards’. The management regimes were as follows: normal management haycut and aftermath grazing (+lime) haycut and aftermath grazing (-lime) grazing only (-lime) grazing only (+lime) hay cut only (-lime) hay cut only (+lime) The results showed that cutting for hay reduced the cover of rye grass and maintained the cover (13%) and number (13) of desirable forb species. Hay cut with aftermath grazing increased the botanical value still further with a mean of 19 forb species on the unlimed and 23 on the limed plots. The findings of this work are useful for managers of moorland edge, though before any change in management, such as incorporating hay cuts and adding lime, is implemented, the current use of the land by other species groups and the implications for them should be considered. Proposed changes should be also evaluated in the context of the surroundings; if the area is dominated by species poor grassland it may be possible to manage a part of it for hay. Oak woodlands The woodland was planted up in the 1990s on a piece of neglected middle hill ground that consisted of rough grass and a small amount of gorse. It is currently not managed and as the trees are growing quite slowly it currently can be described as transitional scrub and gorse. This habitat is very good for passerines. Parc Vicarage wetland This wetland was created by fencing off a wet area that had been part of an intensively managed paddock. It is managed under an ESA tier 2 agreement and is lightly grazed by 16 cattle in spring and late summer. 147 species were identified by a botanist, including 20 lichens and lichenicolous fungi and 15 fungi. Bracken and common dog violet are both present along with some scrub and the site is therefore ideal for dark green fritillary. No further management is required to control the bracken; grazing by cattle should prevent it from spreading. Hafod Grasslands These acid/acid neutral grasslands are part of the Hafod estate that has many designations including SSSI (Site of Special Scientific Interest) and SPA (Special Protection Area). The grasslands, which are grazed by sheep, are dominated by common bent (Agrostis capillaris) and are botanically unexceptional. The site is important however for its colonies of yellow meadow ant (Lasius flavus) and their anthills are numerous here. The anthills can only develop if the disturbance to the sward is minimal; cattle grazing would therefore be inappropriate. The sward needs to be kept short however as the anthills need to be warmed by the sun, if temperatures drop too low, the anthills are abandoned. The site is also important for fungi and 208 species have been identified here including some red data book species. The fungi also require the sward to be short. This is therefore an example of a site that may not initially appear to be of value, but it is clearly important not to attempt to improve the vegetation and sheep grazing at the current levels appears to be the appropriate management regime. Prior to setting new management prescriptions it is important to consider all the taxonomic groups. Ty Coch bracken and scrub This is an area of middle hill that is unimproved and has been only very lightly grazed since the 1960s. Bracken and scrub are therefore increasing in dominance. This is ideal habitat for whinchat for which Wales is a stronghold. It nests under the bracken and uses the fronds as perches. Grazing with cattle, as recommended earlier in this report, would open the sward and increase its suitability for butterflies as well as whinchat. Where bracken is difficult to control and is not suitable for managing for butterflies, it may be possible to plant trees and create woodland. Presence of bracken is often an indicator of an area that was once woodland. Summary From the discussions and comments received during the workshop, it is evident that considerable knowledge gaps remain concerning the management of the moorland edge, particularly with respect to managing the mosaic of habitats present within this transition zone. This mosaic is important for bird and lepidopteran populations, as individual species may use different elements of the mosaic for breeding and feeding. Moorland 17 edge may also offer an important refuge to bird species that are declining in the lowlands. Given these knowledge gaps, the management of moorland edge within existing or proposed agri-environment schemes may need to be reviewed, particularly to determine whether agri-environment prescriptions applied to surrounding land types have any knock-on effects for this transitional zone. For example, does the implementation of reduced stocking rates on heather moorland cause Ffridd land to be utilised more heavily? Moorland edge offers considerable potential for biodiversity conservation and this potential needs to be characterised and quantified. Acknowledgements We would like to thank Tim Milsom, Clare Burrows and Sian Whitehead for their presentations, contributions to the field trip and comments on the report, Sarah Hetherington for chairing the formal session, John Wildig for arranging the field trip, Richard Crompton for talking about the Hafod Estate, Sian Lloyd, Mandy Cowie and Barabara Mclean for help with domestic arrangements and logistical support. Author contacts For further copies of this report, contact Helen Adamson at ADAS Redesdale, Rochester, Otterburn, Newcastle-upon-Tyne, NE19 1SB, or Dr. Sarah Gardner at ADAS Preston, 15 Eastway Business Village, Preston, Lancs, PR2 4WT. References Milsom, T.P., Aegerter, J., Bishop, J.D., Allcock, J.A., Barker, D., Boatman, N.D., Hill, V., Jones, N., Marshall, J., Mckay, N.V., Moore, N.P. & Roberston, P.A. (2003) Review of hill-edge habitats in the uplands of England and Wales. Unpublished report to Defra Conservation Management Division (contract no. BD1235). Parry, M.L., Bruce, A. & Harkness, C.E. (1982). Surveys of moorland and roughland change Series. University of Birmingham, Birmingham, UK. 18