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Photosynthesis Photosynthesis Making the fuel of life Sunlight energy is trapped and converted into chemical energy in the form of glucose Energy is stored in organic molecules made of inorganic raw materials Photosynthesis is considered and anabolic process: Involves the synthesis of new, larger compounds Organisms that are able to make their own food in this way are called autotrophs Plants, Algae, and Cyanobacteria Specific organelle (Eukaryotes): Chloroplast Photosynthesis Making the fuel of life What is used? What is produced? + 6CO2 + 6H2O C6H12O6 + 6O2 Glucose autotrophs What living things make photosynthesis? Algae (seaweeds) and plants: The entire process takes place in the chloroplast Kelp Some unicellular Eukaryotes (Phytoplankton) make photosynthesis, and some bacteria (Cyanobacteria, that lack chloroplasts) Anabaena, a cyanobacteria Mosses, ferns, pine trees, and flowering plants Euglena, inside the group Protista, is an unicellular Eukaryote How much is produced? 160,000,000,000 tons per year… or… 352,000,000,000,000 pounds 1 Photosynthesis Why are these living things autotrophs? •They have chlorophyll •A green pigment that absorbs light energy and initiates the sequence of reactions (Able to produce their own food) Chlorophyll Is contained folded membranous structures called thylakoids in chloroplasts Sunlight provides the energy Chlorophyll molecules absorb photons (units of energy) Electrons in the covalent bonds of the chlorophyll gain energy With this energy, electrons are able of entering into chemical reactions responsible for the production of ATP + NADPH ATP + NADPH are used in turn to produce glucose Why is chlorophyll green? •Light consists of high-speed particles called photons. •Photons are packets of electromagnetic energy Electromagnetic spectrum shows the full range of radiation types Chlorophylls (a and b) Absorb mostly blue and red light But reflect 100% of the green light 2 Photosynthesis Why do color of leaves turn to different colors in fall? Why are there brown and red algae? Other pigments absorb slightly different wavelengths and “help” the chlorophyll! Carotenoids: in plants and algae Phycobilins: in algae and bacteria Chlorophyll c & d: in algae Why having accessory pigments? With only chlorophyll Visible spectrum With accessory pigments Visible spectrum Ergo… The more pigments The more “types” (different wavelengths) of energy autotrophs can absorb Stages of photosynthesis (1) Light dependent reactions Need light to happen Take place in the membrane of the thylacoids The light reactions convert solar energy to chemical energy (ATP, NADPH) (2) Light independent reactions or “Dark”reactions or “Calvin cycle” DOES NOT need light to happen Takes place in the stroma (matrix) of the chloroplast Uses the products of the light reactions to make sugar from carbon dioxide 3 Photosynthesis How do plants exchange the gases involved in photosynthesis? Stomata are tiny pores in leaves where carbon dioxide enters and oxygen exits. Produced in the stroma of chloroplast “in” using stomata “in” using roots “in” and “out” using stomata (1) Light dependent reactions How chlorophyll traps sunlight and passed the energy to the ETC… Chlorophyll is arranged in plants in photosystems Which are molecular light-capturing devices Two sequential photosystems are needed to complete the light reactions Why splitting water? e- (electrons) released are used to replace those lost by the chlorophyll H+ and O2 are also produced ETC ETC NADPH production ATP production Water-splitting photosystem NADPH-producing photosystem Both are used in the dark reactions to produce glucose 4 Photosynthesis (1) The “excited” electrons are passed through two electron transport chains (2) This results, as in cellular respiration, in H+ being pumped into the cavity of the thylacoid… and electron “jumping” from one acceptor molecule to another, with each “jump” meaning higher energy levels (3) Also as in cellular respiration, H+ return to the exterior of the thylacoid down a gradient concentration and ATP is generated (2) “Dark” reactions or Calvin Cycle Input Functions like a sugar factory within the stroma of a chloroplast enzyme Three CO2 CARBON FIXATION Regenerates the starting material with each turn Six Three ADP Three Six ADP + Six Calvin cycle Six Six NADP ATP (energy) and NADPH ( H+) generated in the previous stage, atmospheric CO2, and a starter sugar molecule allow to build organic molecules (ex:glucose) One G3P Output Glucose and other compounds 5 Photosynthesis The first real product of photosynthesis is Glyceraldehyde 3-P instead of glucose! This molecule is the main intermediate product of glycolisis in cellular respiration The surplus of Glyceraldehyde 3-P is used to build glucose and a diversity of molecules How do plants adapt to water scarcity? Plants open their stomata in the epidermis to take in the CO2, but H2O may be lost by dehydration CO2 CARBON FIXATION Calvin Cycle enzyme “C3” photosynthesis Sugar (most plants) The enzyme that “fixes” the CO2 works very well when CO2 is abundant, but poorly when CO2 concentration is low “C4” photosynthesis “CAM” photosynthesis (Hot and dry areas) (also in hot and dry areas) stomata slightly open in the epidermis to take in the CO2, this minimizes water loss stomata open at night to take in the CO2 CO2 CO2 C CARBON FIXATION uptake Different enzyme, very Calvin Cycle Sugar Night efficient when CO2 concentration is low (when stomata are slightly open Calvin Cycle But there are extra steps needed and more ATP is used… more energy is spent C uptake Day CARBON FIXATION stomata closed during the day, this minimizes water loss Sugar 6