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Environments Through Time 1 Evidence from early Earth indicates the first life forms survived in changing habitats during the Archean and Proterozoic eons identify that geological time is divided into eons on the basis of fossil evidence of different life forms define cyanobacteria as simple photosynthetic organisms and examine the fossil evidence of cyanobacteria in Australia outline the processes and environmental conditions involved in the deposition of a Banded Iron Formation (BIF) examine and explain processes involved in fossil formation and the range of fossil types outline stable isotope evidence for the first presence of life in 3.8 x 109 year-old rocks gather and process information from secondary sources to draw up a timeline to compare the relative lengths of the Hadean, Archaean, Proterozoic and Phanerozoic eons gather and analyse information from secondary sources to explain the significance of the Banded Iron Formations as evidence of life in primitive oceans gather, analyse and present information from secondary sources on the habitat of modern stromatolites and use available evidence to propose possible reasons for their reduced abundance and distribution in comparison with ancient stromatolites Geological time and eons: - The geological time scale is divided into 4 eons: Hadean, Archaean, Proterozoic and Phanerozoic. They are divided on the basis of fossil evidence of different life forms. Hadean: (4.5-4 b.y.a) no rocks have survived from this period. Zircon (pieces of original rock have evidence of life) - Archaean: (4-2.5 b.y.a) granitoids; greenstones; basalts and pillow rocks (only formed in the oceans therefore oceans must have existed). All the rocks were highly metamorphosed. Cores of continents formed during this eon. Very ancient life forms (trace fossils only) - Proterozoic: (2.5 b.y.a-545 ma) plate tectonic super cycle established. Sedimentary rocks deposited on crust (sandstone, conglomerate, mudstone, limestone). Fossil bacteria and algae - Phanerozoic: (545 ma –present) Complex fossils with hard parts appear. Lots of material is added to continents at subduction zones Cyanobacteria, stromatolites and banded iron formations - Cyanobacteria were the first organism to use photosynthesis as a means to respiration Cyanobacteria are simple photosynthetic organisms Sunlight/chlorophil Carbon dioxide + water --------------------------- Glucose + oxygen Cyanobacteria and Fossils: - The remains of cyanobacteria can found as stromatolites in Western Australia The North Pole district in Western Australia is home to many stromatolite fossils Stromatelites found in Marble Bar W.A in the Pilbura Region are the oldest fossil evidence for the presence of cyanobacteria 3.5 BYA a stromatelite is a rocky layered deposit formed by communities of Cyanobacteria Modern Stromatolites require: - shelter from predators warm water hyper saline shallow water low amounts of tidal flushing form domes, mats, columns or cones Conditions which stromatolites needed: - Required protection from UV light, a few metres under water was suitable warm; shallow; hypersaline water, shelter from predators and low amounts of tidal flushing. - Stromatolites have declined over millions of years this is due to an increase in oxygen concentration , this would reduce the population of cyanobacteria Banded iron formations and chemical precipitation: - Banded iron formations are sedimentary deposits of banded iron oxide which are composed in alternating layers Formed through chemically precipitated iron oxide Most Banded iron formations were deposited 2 – 2.5 bya Chemical Precipitation: - Photosynthetic cyanobacteria produced oxygen and oxygenated the upper layers of the ocean. This oxygen caused chemical precipitation The ocean was rich in dissolved in iron, oxygen reacted with this iron to create iron oxide, which precipitated to the bottom of the sea floor and formed alternate layers The formation of banded iron moderated and controlled the amount to oxygen in the atmosphere, when all the dissolved iron was precipitated there was an explosion of oxygen in the atmosphere Discuss the significance of BIF’s as evidence of life in primitive oceans: - BIFs provide evidence of primitive life in the oceans by the presence of oxygen that must have been caused by photosynthesis by life forms. Fossil formation and fossil types - Fossils are the preserved remains or impressions of prehistoric life forms or evidence for their existence - MOULD - the shape/imprint of the organism CAST - the imprint filled in with new minerals Conditions required for fossil formation: (BSCA) 1. Burial (rapid) 2. Sediment (continual supply of sediment to avoid erosion) 3. Compaction (of sediment) 4. Anoxic conditions - Fossil formation is unlikely for a number of reasons o Many deceased organisms are eaten by scavengers or broken down by microbes o The conditions for burial are not met ie rapid burial, continual supply of sediment, compaction and anoxic conditions Fossil forms: Unaltered soft parts – When bacteria and mould are prevented from acting of soft tissue Unaltered hard parts – Soft tissue is decomposed, however, hard parts remain eg bones Altered hard parts – When fossils undergo physical and chemical changes. Eg hard parts replaced by with silica, pyrite or iron oxide. Trace fossils – Fossils leave a mould which is filled in by minerals creating a cast. These can be in the forms of imprints of organisms and burrow Fossil types: - Examples o Unaltered soft parts Mammoths in Siberia Animals trapped in tar pits o Unaltered hard parts Shells of mollusks Hard parts of insects in tree resin o Altered hard parts Petrified wood Carbonisation where a fossil turns to black carbon o Trace fossils very common impressions and moulds Casts of original organisms Footprints of organisms eg dinosaurs Evidence for first life: - The earliest evidence of life is found in metamorphosed sedimentary rock 3.8 b.y.o found in ISUA, WEST GREENLAND. The graphite minerals enriched in Carbon 12 in these rocks provides a “signature” of life. There is a high Carbon 12: Carbon 13 ratio 2 The environment of the Phanerozoic eon outline the chemical relationship between ozone and oxygen explain the relationship between changes in oxygen concentrations and the development of the ozone layer analyse information from secondary sources to identify the major era subdivisions used to describe the Phanerozoic and describe the general differences in life forms in each era describe the role of ozone in filtering ultraviolet radiation and the importance of this for life that developed during the Phanerozoic eon Chemical Relationship between ozone and oxygen: - Ozone is formed in the stratosphere (upper atmosphere Ozone is formed when UV rays separate 02 (oxygen) into separate 0 molecules As 0 atoms are extremely unstable they are forced to react with a nearly atoms usually other 0 atoms or 02 atoms If they react with 0 atoms then 02 will be formed, however, if reacted with 02then 03 (Ozone) will form 0 + 02 03 Changing oxygen concentrations and development of ozone layer The presence of oxygen in the atmosphere: – increased ozone production and therefore reduced UV radiation that reached earth – This allowed for organisms to evolve into more complex life reducing the UV damage emitted by the sun – increased oxygen lead to global cooling (previously the Archaean was very warm as it had high CO2 levels) Role of Ozone and importance: - the introduction of ozone layer meant that life could evolve beyond the deep areas of the ocean ozone is able to filter and prevent most of the dangerous UV radiation from reaching earth therefore moderating and controlling global temperature This is important as UV radiation is very damaging. Terrestrial organisms need ozone in order to prevent cellular death by disrupting DNA, mutations (cancer), cataracts (in eyes), sunburn, and death of some organisms Identify the major era subdivisions used to describe the general differences in life forms in each era: The Phanerozoic Eon Eon Era Life Forms Cenozoic (144 ma-present) -mega fauna extinction Phanerozoic -mammals, birds and flowering plants dominate -homo sapiens Mesozoic 248-144 ma -flowering plants -land organisms dominate -reptiles and dinosaurs Paleozoic 540-248 ma -first jawed fish - vertebrates -amphibians -land plants 3 The Cambrian event interpret the relative age of a fossil from a stratigraphic sequence compare uses of relative and absolute dating methods in determining sequences in the evolution of life forms distinguish between relative and absolute dating discuss the possible importance of the development of hardened body parts in explaining the apparent explosion of life in the Cambrian period process information to examine at least one example of a stratigraphic sequence and describe any fossils found in this sequence use available evidence, including computer simulations, models and photographs to examine the changes in life forms that occurred during what is commonly referred to as the ‘Cambrian event’ deduce possible advantages that hard shells and armouring would have given these life-forms in comparison with the soft-bodied Ediacara metazoans of the late Proterozoic, in terms of predation, protection and defence - - The Cambrian Event The Cambrian explosion/event was the almost immediate radiation of shelled organisms during the early Cambrian The Cambrian Period marks the boundary between the Proterozoic and Phanerozoic eons and the almost instantaneous appearance of a variety of shelly animal fossils which are ancestors of present day life e.g. the Pikara Graciens At this boundary there is a distinct transition between soft-bodied organisms to hard shelled creatures 50 ma after the Cambrian boundary there is an increase in complexity of fossils possible events that caused the Cambrian explosion o Increased oxygen supply allowed larger organisms to survive o Predation allowed organisms to obtain more nutritious food supply (theory not proven) o Genetic mutations increased the rate of evolution (theory not proven) Distinguish between relative and absolute dating - Relative dating is a type of dating, whereby the rock is compared to other rock layers. Ie rock layer 2 is older then layer 3 but younger then layer 1 Law of superposition, states that the rock layer on top of the previous rock layer is older. (see below) - Absolute dating determines the age of specimens Absolute dating is achieved through radiometric dating Components of atoms o Neutrons (n) o Protons (p) o Electrons (e) Compare uses of relative and absolute dating methods in determining sequences in the evolution of life forms Relative dating refers to determining rock/fossil age in reference to other rock layers and the geological time scale. It does not classify a rock strata in terms of years. Absolute dating - determining the exact age of a rock/fossil in years - The rock must be igneous - Radioisotope dating is based on the decay of radio isotopes found in the rocks to determine the fossil/rocks age in terms of years. Half lives are used to determine the age of breakdown from when the parent rock first formed. Half Life, the half life of an isotope is how long it takes for half of the parent isotope to decay into the daughter isotope. If the rate of decay of isotopes is known then it is possible to compare the amount of parents isotopes present with the amount of daughter isotopes present to determine age Example: o Potassium – 40 -> Argon – 40 o Carbon – 14 -> nitrogen - 14 Discuss the possible importance of the development of hardened body parts in explaining the apparent explosion of life in the Cambrian Period There are 2 main points why hardened body parts explain the Cambrian explosion: 1. Hard parts provided support for muscle joints and internal organs in organisms and offered protection from predators and harmful UV rays allowing these organisms to better survive and move onto life where food availability was higher 2. Hard parts are easily fossilized unlike soft parts leading to an increase in fossils with hard parts found so a sudden explosion of life seems to occur during the Cambrian Period. Deduce possible advantages that hard shells and armoring would have given these life-forms in comparison with the soft bodied Ediacara metazoans of the late Proterozoic, in terms of predation, protection and defense – Hard parts provided animals with o protection from predators o a skeleton to provide a framework to support feeding organs of filter feeders and suspension feeders o skeletons allow organisms to grow larger and therefore more protection from predators o skeletons also provide firm points for muscle attachment to improve mobility o protection from UV o stops organisms in intertidal zones from drying out during low tides - The Cambrian animals had an advantage over the Ediacara Metazoans as they had hard parts and were more mobile which allowed them to escape from their predators and move to areas where more food was available. 4 Exploiting new environments outline the theory of evolution by natural selection outline evidence that present-day organisms have developed from different organisms in the distant past gather and analyse information from a geological time scale and secondary sources to identify and date the major evolutionary advances made by plants and animals summarise the main evolutionary changes resulting from the selection of living things exhibiting features that allowed them to survive in terrestrial environments gather information from secondary sources to summarise the features and distribution of some of the first land plants, amphibians and reptiles outline the major steps in the expansion to the terrestrial environments by land plants, amphibians and reptiles gather information from secondary sources to compare the diversity and numbers of organisms from a fossil site identify the advantages the terrestrial environment offered the first land plants and animals Outline the theory of evolution by natural selection - Organisms that are best suited to the environmental conditions will survive. These organisms will multiply producing offspring with their favorable characteristics/genes. They will dominate the population and the others will die out. Outline evidence that present-day organisms have developed from different organisms in the distant past - the fossil record shows the gradual progression of simple life forms into more complex organisms - Comparative embryology o The comparison of embryos in organisms - Anatomy o The structure of an organism such as the pentadactyl limb found in a variety of organism including humans, horses, lizards, cats and birds - Biogeography o Where similar things live and their habitats o For example the golapocus islands o The crimson rosella and the eastern rosella - Physiology o The functioning of an organism o Eg Hemoglobin - Biochemistry o The analysis of DNA and Proteins. o Many organisms have similar cell structure and function. o The more similar the DNA and proteins in organisms the more closely related they are o DNA analysis has become the most sophisticated and used means to determine evolution and similaritie TGA fly TGA Frog TAG Mouse GGG cockatoo DNA sequencing TGA common ancestral sequence Summaries the main evolutionary changes resulting from the selection of living things exhibiting features that allowed them to survive in terrestrial environments - - - - Water o Plants developed roots to ground themselves into the soil and absorb water o Plants also developed an epidermis which reduced the amount of water lost o Animals developed more complex nervous systems which would allow for animals to sense and locate water o Animals developed skin which reduced the amount of water lost Oxygen o Plants developed stomates which stopped water imbalances through respiration o Plants created oxygen the process of photosynthesis o Animals developed lungs which allowed for aerobic respiration Reproduction o Evolved seeds which were able to spread via the wind, water and by other animals thus transporting offspring over large areas o Animals developed internal fertilization Energy o Plants developed a supportive structure which allowed them to be closer to the sun and as such photosynthesis would occur easier o Animals waited until plants moved to the terrestrial environment as plants were an energy source for animals Identify the advantages the terrestrial environment offered the first land plants and animals Advantages Disadvantages - more abundant O2 - direct exposure to air=drying out therefore animals developed scaly skin - more light for photosynthesis - new mode of fertilization needed which lead to the development of internal fertilization and hardshelled eggs for protection - more available habitats - lung development needed (evolution) - less competition for food - gravity force is greater on land and therefore skeletons (fauna) and lignin (plants) were developed to hold against gravity Outline the major steps in the expansion to the terrestrial environments by land plants, amphibians and reptiles - All life was marine based until the Ordovician. Early plants colonized near shore environments which encouraged animals to move closer to the food supply 1) Plants 2) Insects 3) Animals - - - - - - - Precambrian o Cyanobacteria o Metazoans Cambrian o Anthropods o Brachiopods Ordovician o Fish Silurian o First vascular land plants – liverworts o First land animals Devonian o Amphibians o Land plants Carboniferous o Reptiles Permian o Mammal like reptiles o Ammonites Triassic o First mammals o Dinosaurs Jurassic o Flowering plants Tertiary o Hominoids Lobe Jawless bony – plated fish Fish with Jaws Bony Fish Cartilaginous fish Early amphibians Modern amphibians Primitive reptiles Modern birds Modern reptiles Mammals 5 Past extinction and mass extinction events compare models of explosive and gradual adaptations and radiations of new genera and species following mass extinction events distinguish between mass extinctions and smaller extinctions explain the recent extinction of the marsupial, bird and reptile megafauna in Australia, as an example of smaller extinction events involving several large species. compare these smaller extinction events with widespread ‘catastrophic’ events in which entire ecosystems collapse with the extinction of many entire classes and orders gather, analyse and present information from secondary sources to compare two different concepts used to explain mass extinction events gather information from secondary sources and use available evidence to identify the relationship between mass extinctions and the divisions of the geological time scale analyse information from secondary sources on at least two different hypotheses used to explain the extinction of the megafauna assess a variety of hypotheses proposed for the mass extinctions at the end of the Permian and at the end of the Cretaceous Compare models of explosive and gradual adaptations and radiation of new genera and species following mass extinction events - Radiation: o occurs after extinctions when the remaining groups of organisms undergo rapid evolutionary expansion to replace the lost organisms. o The new genera or species that radiates have similar features to their ancestors but are specifically adapted to their environment. o After this, rates of evolution slow dramatically due to increased competition for food/shelter and predators therefore only new genera or species evolve opposed to phyla and classes. - The geological timescale indicates that there is a constant cycle of extinction and radiation. The fossil record indicates that evolution has occurred gradually and quickly (in geological terms) Gradual: When evolution occurs over a long period of time through continuous changes in Explosive: punctuated equilibrium theory proposed by Eldridge and Gould suggests that evolution occurs by rapid bursts of change within an organism followed by long periods of stability Past extinction and mass extinction events Extinction: when species die out. They occur due to meteorites, volcanoes, regression/transgression. Radiation: when the survivors of the extinction radiate and flourish replacing the lost organisms Ecological Niche: the place of a plant or an animal within its ecosystem or community Distinguish between mass extinctions and smaller extinctions - - Mass extinctions o Affect a variety of species over a large area potentially globally or in most areas of the world o Typically relatively brief and followed by explosive radiation bringing about new species and a new genetic variety o Such as the K/T Cretaceous and Permian extinction Smaller extinctions o Affect only a relatively small amount of species in a relatively small area o Such as extinction of Australian megafauna Explain the recent extinction of the marsupial, bird and reptile mega fauna in Australia, as an example of smaller extinction events involving several large species. - The Australian megafauna extinction is considered a small extinction as it occurred in one particular geographical area and a number of species managed to survive There are two theories to explain the extinction, it could have been a combination of the two - Climate change o occurred 18 000-22 000 years ago o Australia’s climate became more arid as it drifted northward- changed from cold and dry to warm and dry o ecosystems changed from forest to grassland and surface water became scarce o many organisms lost their habitats and these pressures were too extreme to adapt quickly enough o Smaller species managed to survive as they are in need of less food and are able to reproduce more offspring - Human intervention o Aboriginal human intervention at fault o mega fauna’s large size made them slow making them vulnerable to human predation o human arrival coincides with the beginning of the extinction o Some evidence of attacks on megafauna found in fossil remains of bones o humans also induced vegetation change such as back burning o unable to adapt to the introduction of human predation Compare these smaller extinction events with widespread ‘catastrophic’ events in which entire ecosystems collapse with the extinction of many entire classes and orders - The Permian extinction occurred at the end of the Permian The extinction caused the death of almost 95% of species on Earth The extinction was on a global scale affecting all species The extinction was either caused by a catastrophic event ie a bolite collision, a gradual event such as sea level changes or a combination of both Assess a variety of hypotheses proposed for the mass extinctions at the end of the Permian and at the end of the Cretaceous Catastropic: Cretaceous K/T - - Suggests a bolide (meteorite ) hit earth killing all organisms o The bolite may have triggered large tsunamis o Caused bushfires on a large scale o Caused death to organisms in immediate vicinity o The resulting ash cloud may have caused a global cooling affect Evidence for bolide impact: o Impact crator (chixulub crator) Located in the golf of Mexico 10km radius dated back 65 million years Iridium levels are high at the KT boundary layer. Clay in these rocks contain unusually high iridium which are typically found in meteors There are three unusal deposits found at the KT boundary Tsunami deposits Impact debris (tektites) Fullereness in boundary clay indicating high temperature and combustion Gradual: Permian - - Transgression/regression o A sudden drop in sea level globally causing geologically unstable oceans and placing immense stress on marine organisms especially since most organisms at the time were in shallow areas o Later the sea level rose, again placing stress on species, when they had already adapted to the previous ocean levels o This may have also affected rain on a global scale altering rainfall and altering climatic conditions Volcanic pulses o Theory suggests that Volcanic eruptions lasting up to 10,000 caused a the cooling of the Earth by blocking the sun The ash then settled and CO2 caused global warming Oceans may have then warmed and became anoxic