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STEPPE COLLECTION Panayoti Kelaidis, Senior Curator and Director of Outreach; Leesly Leon, Docent Coordinator, November 2013 Description Plants in the collection come from the steppe biome which is a dry, cold, grassland found in all of the continents except Australia and Antarctica. It is mostly found in the USA, Mongolia, Siberia, Tibet and China. Steppe regions are located away from the ocean and close to mountain barriers and therefore have low humidity. The collection is divided in the following sub-collections: • • • • Asian Steppe: plants which are naturally occurring in the vast central region of the Asian continent. South American Steppe: plants which are naturally occurring in very small areas of South America (very few in our collection). African Steppe: plants which are naturally occurring in very small areas of Africa, predominantly South Africa (few in collection). North American Steppe: plants which are naturally occurring in a large central area of North America. History of the Collection The Steppe Collection finds its origins in the Rock Alpine Garden where, up until the advent of the WaterSmart Garden, it was the only garden where visitors could find exogenous (non-native) steppe plants within Denver Botanic Gardens. It was here where successful steppe plants were tested, while the concept of the Watersmart® Garden, in 1994, led to the increase in use of Asian and Southern Hemisphere steppe plants. Native steppe collections also found expression in the Plains and Dryland Mesa Gardens, developed in the late 1980’s. The Plant Asia Garden designed by Gardens staff and constructed in 2001-2002 includes a section where drought tolerant plants from the Asian steppe are showcased. These materials are for Denver Botanic Gardens use only. STEPPE COLLECTION 1 Relevance There are four major areas of the world with a very similar climate to that of the North American steppe, including short grass and shrub steppe in Colorado. The high shrublands and grasslands of the steppe regions of the world share Denver’s temperate, semi-arid climate. Many beautiful and unusual Asian, African and South American plants are ideal candidates for a Rocky Mountain garden. Since steppe plants have evolved in climates with intense sun year around, they often have much more persistent, wintergreen foliage that adds color to our long “dormant” season. Collection Facts • • Both the high, dry steppes of Asia and the high, dry steppes of N. America (which is where Denver is located) are located on a plateau and have many of the same genera. The steppe collection is the only one of its kind in North America, making a visit to Denver Botanic Gardens unique. Steppe Climate Facts • • • Steppe areas are defined as semi-arid grasslands that receive 10-20” of rain each year. Steppe areas are treeless and make up 15% of the Earth’s surface. The term steppe comes from the Russian word which means flat, arid land and denotes the climate too dry to support a forest but not dry enough to be a desert. These materials are for Denver Botanic Gardens use only. STEPPE COLLECTION 2 WATER-SMART GARDEN® Silver Horehound • • • • • Scientific name: Marrubium incanum Family: Lamiaceae (Mint) Native range: Italy, Sicily, Balkans Marrubium is a genus of some 40 species of perennials from the Mediterranean Europe and temperate Asia which are predominately found in sunny, rocky, and dry wasteland. Their two lipped flowers are borne in auxiliary whorls where they bloom during the summer season. Variegated Statice • • • • • • • • Scientific name: Bukiniczia cabulica Family: Plumbaginaceae (Leadwort) Native range: Pakistan, Afghanistan The blue-green leaves with white veins form a rosette up to 6 ± inches in diameter. A biennial, producing a 14 ± inch inflorescence with tiny nondescript whitish/pink flowers in its second year. A prolific seeder - within a few years one is likely to see baby B. cabulicas all about the garden. Good plant for rock gardens. The specific name, cabulica suggests a possible tie to the Kabul, Afghanistan region. Skullcap • • • • • • • • Scientific name: Scuttelaria orientalis Family: Lamiaceae (Mint) Native range: Russia Genus is widespread in temperate regions and on tropical mountains. Most are annual or perennial herbaceous plants from 5 cm to 1 m tall, but a few are subshrubs (a woody perennial plant - distinguished from a shrub by its ground-hugging stems and lower height). Photo by Panayoti Kelaidis They have square stems and opposite leaves (characteristic of the mint family). The flowers have upper and lower lips. The genus is most easily recognized by the typical shield on the calyx that has also prompted its common name. These materials are for Denver Botanic Gardens use only. STEPPE COLLECTION 3 PLANTASIA GARDEN Turkish Veronica • • • • • • • • • • • Scientific name: Veronica liwanensis Family: Scrophulariaceae (Figwort) Native range: Turkey 1997 Plant Select® winner. Sturdy groundcover with waxy, succulent - like, teardrop - shaped leaves. Evergreen Xeriscape, perennial plant. Cobalt blue flowers (blooms April - June in Colorado). Veronica liwanensis can be used like thyme to create a ground cover “lawn.” With a little extra water, Veronica liwanensis re-blooms lightly later in summer. Needs afternoon shade in hot summer climates. Great ground cover at the base of evergreens where it still gets some sun. Iberis taurica • • • • • Family: Brassicaceae (Mustard) Native range: Asia Minor Flowers are white/lilac tinged. Forms cushions of rubbery, dark burgundy-green leaves. Annual or biennial. Tulip ‘Red Riding Hood’ • • • • • • • • Scientific name: Tulipa greigii ‘Red Riding Hood’ Family: Liliaceae (Lily) Native range: Cultivated Tulipa greigii ‘Red Riding Hood’ has very bright red flowers with a black heart. Foliage is streaked with purple stripes. Low growing so they should be planted in the front of a garden bed. Easily grown in average, medium, well-drained soil in full sun. Good for rock gardens and beds, border fronts or around trees or shrubs. www.mobot.org Vvendensky's Tulip; Tulip • • • • • Scientific name: Tulipa vvedenskyi 'Tangerine Beauty' Family: Liliaceae (Lily) Native range: Central Asia Striking tulip species with bright scarlet-orange flowers. One of the hallmarks of this species is the wonderful undulate foliage. These materials are for Denver Botanic Gardens use only. STEPPE COLLECTION 4 Turkestan Onion • • • • • • • • Scientific name: Allium karataviense Family: Amaryllidaceae (Amaryllis) Native range: Kyrgyzstan, Tajikistan, Uzbekistan Found on mountain slopes. Sunny places but it also tolerates shaded locations. The leaves are tongue shaped and green with a width of 10 cm and a length of 30 cm. Flowers are growing on a central stem with a height of 30 cm. Up to 40 flowers can be found on one stem. Color can very between pink and white (the color gets paler with time). Photo by Anna Russo Spike Thrift or Prickly Dianthus • • • • • • • Scientific name: Acantholimon hohenackeri Family: Plumbaginaceae (Leadwort) Native range: Caucasus, Iran Mounded blue-gray foliage with very prickly needle-like leaves Pink flowers. Prefers well-drained soil. Acantholimons were used as a source of fuel. Persian Stonecress • • • • Scientific name: Aethionema grandiflorum Family: Brassicaceae (Mustard) Native range: Iran, Iraq, Caucasia A small genus of plants native to the Mediterranean, Europe and southwest Asia with flowers in crowded terminal racemes - a flower cluster inflorescence in which the flowers are borne on short stalks along a long main stem. Narrow-leaved Foxtail Lily • • • • • • • Scientific name: Eremurus stenophyllus Family: Xanthorrhoeaceae (Grasstree) Native range: Central Asia to W. Pakistan Spikes of dense 4 cm golden yellow, star-shaped flowers begin opening at the bottom of the spike first and slowly bloom to the top. Like shelter and warmth, especially after flowering. The leaves appear early in wide clumps, and start to die down just as the flower spikes appear. The foxtail lily is hardy and prefers a sandy, well-drained soil. These materials are for Denver Botanic Gardens use only. Photo by Panayoti Kelaidis STEPPE COLLECTION 5 ROCK ALPINE GARDEN Fringed Wormwood or Fringed Sage • • • • • • • • • • • • Scientific name: Artemisia frigida Family: Asteraceae (Aster) Native range: SE & E. Russian Federation, N. America Perennial. The flowers are hermaphrodite (have both male and female organs) and are pollinated by wind. Prefers light (sandy) and medium (loamy) soils, requires well-drained soil and can grow in nutritionally poor soil. The plant prefers acid, neutral and basic (alkaline) soils. It cannot grow in the shade. It requires dry or moist soil and can tolerate drought. Photo by Anna Russo frigida = cold; rigida = rigid The leaves were used by the Hopi Indians as a flavoring for sweet corn. An infusion of the leaves is used in the treatment of indigestion; coughs and colds (chew the leaves and drink the juice to treat heartburn). Leaves are also placed in the nose to stop nosebleeds. Tea made from the leaves is used to relieve menstrual pain and discomfort. Both the growing and the dried plant can be used as an insect repellent. Salvia pisidica (on the moraine mound) • • • • • • • Family: Lamiaceae (Mint) Native range: Turkey In Turkey it grows on dry limestone slopes. Here it needs a very well-drained soil, good for rock gardens. Maximum height is about 9 inches, with a spread of up to 2 ft. Grey-green leaves contrast well with the striped blue and white flowers. It can be propagated by cuttings which can be slow to root, or from seed which can be slow to germinate. Stipa ucrainica (in the lower meadow) • • • Family: Poaceae (Grass) Native range: S. Russia, Turkmenistan Displays beautiful seed heads These materials are for Denver Botanic Gardens use only. STEPPE COLLECTION 6 SOUTH AFRICAN PLAZA GARDEN Berkheya montana • • • Family: Asteraceae (Aster) Native Range: South Africa The plant looks a thistle – some consider it as such – but it displays a beautiful flower Ruschia putterillii • • • • • Family: Aizoaceae (Fig-Marigold) Native range: South Africa Suitable for growing in containers Drought tolerant It looks like an ice plant but it is shrubby Photo by Panayoti Kelaidis Kangaroo Grass • • • • • • Scientific name: Themeda triandra Family: Poaceae (Grass) Front range: Old World Tropics & Subtropics: [ native to India, Korea, China, and Japan (1849)] This is a lovely green to blue-green, tufted grass that is often flushed with pink and turns red with age. Some forms have bright yellow culms (stems). Plants from higher altitudes tend to be shorter and dark purple, whereas, at lower altitudes, plants are often lighter coloured and flushed only with purple. Seeds for the plant in this garden was collected by Dan Johnson, Curator of Native Plants at DBG These materials are for Denver Botanic Gardens use only. STEPPE COLLECTION 7