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BLUMEA (1977) 23 337—340 Pacific capsular Myrtaceae Tristania (New Caledonia) Dawson J.W. Botany Department, of 12. Wellington, Victoria University Zealand. New INTRODUCTION Brown, Aiton’s Hortus Kewensis (2nd Ed.) Tristania R. with three species have since been added three original species Caledonian species of the groups selected the become same to at grow it. Six established was species are the generic at decided which yet been group. It has but ifthe T. laurina not currently recognised on in are groups of the New name, low elevations in scrub and forest is group (1863) 10 not would 371, Caledonia where New ultrabasic rocks. Species of the Guinea, Borneo, and New these level. All Gris, Bull. Soc. Bot. Fr. et has shown that the and further that groups original generic found in Australia, are 417, study (Wilson, 1971) recent the Tristania laurina Tristaniopsis Brongniart name group to a their separation warrant belong and three different to should retain the available for they mostly genus belong different sufficiently the to (1812) 4 T. laurina, and T. conferta. A number of other species neriifolia, T. — probably elsewhere in Malesia. DESCRIPTION Shrubs smooth; nate or branching trees; bud scales with a a TRISTANIA monopodial; leaves wanting; 2/5 phyllotaxy, OF more (NEW CALEDONIA) or less dark, stems young leptophyllous to older microphyllous, stems paler and pronounced ridge extending from below each leaf insertion through several internodes; young parts densely clothed with short white reddish or simple to hairs, older parts glabrous compound dichasial branches each terminated by subtend (figs. or sometimes as with ovary, brescent or orbicular, style, to stamens to reduced to node (fig. bud; bracts adnate except triangular, stamens (figs. distal parts of acute; 6, half their lengths, with median stamens the 7) in 2—70 stamens about as long tip of the the locules (fig. 6) or ovary; more stigma small, to or 9, acute and style, 5, or more the in fascicle, long as less petals, one to the petals, small, dorsifixed, versatile, Ovary (figs. as 10), 6, the longest placentas apical less axile (T. glauca, figs. as on they petal surfaces gla- per twice connective. convex, inner fascicles opposite 5 superior, three-locular; style (figs. 6,9) about as long into the top of the axes stamens petals (fig. 5) much shorter; anthers (figs. 8, 9) at the to i); bracts and bracteoles lanceolate, pubescent 5, long, in upper leaf axils single flower, a hypanthium, filaments, and white; single prominent oil gland superior set up or outermost the lateral and inner next all parts sepals (fig. 4) yellow, orange, several series, the a the as or remaining pubescent. Inflorescences (fig-*) dormant vegetative inner surface glabrous; rarely single, fused for with far 3). Flowers either with 2, or cymes a or alter- dorsiventral, 9, in the inner in the latter 10) semi- stamen, not angles case a of stylar 338 BLUMEA VOL. 23, No. 2, 1977 J. Figs, size — 1 T. callobuxus. 8. T. X15. Petal; X5. 11. X15. 17. — Fertile T. one Cell locule; X15. T.S. X5. Embryo; X15. — 19. Sterile — 23. T. callobuxus: Placenta T.S. 14. — and seed; X5. — capitulata. WELTU (fig. T. ovule L. group S. L. S. detail T.S. T. glauca. T.S. inner T. into X10. 9634 — 10. stylar tip of ventral is 16. seeds stippled; X5. nat. 5. L. S. just ovary; above — pla- Dehisced — 18. T. 15. T. capsule; capitulata. fertile seed, pigmented cells stippled; X300. seed; X175. (Tristania glauca: (figs. 8, 9), guillainii: McKee 28014). McKee — 22. McKee 20201, 25585 T. — T. glauca. Ovule 13. T. callobuxus. — view; X15. T. glauca. canal — branch; Sepal; X5. integument stippled; X50. — Cell detail T.S. testa sterile cotyledons; Anther, locule; X15. one at 4. T. callobuxus. flower; X5. of fertile testa vegetative bud — Opening from ovule, capsule, wings 12230, WELTU 19); glauca. 339 12 callobuxus. 7. T. ovules removed. guillainii. Cell 21. 9. — integument stippled; X300. undehisced capitulata. 29143, McKee 28863 T. inner — with Dormant Bracteole; X3. L. S. flower; X5. view; X15. ovule, T. T. 3. T. callobuxus. into locule T. callobuxus. detail capitulata. McKee dorsal Pacific Capsular Myrtaccae T. glauca. Inflorescence. 1. — T. callobuxus. 6. T. callobuxus. seed; (fig. 21); — Anther, 12. — Bract; X3. glauca. View T. — from group capitulata. 20. callobuxus. callobuxus. — centa; X3. Tristania R. Br. —23. 2. W. Dawson: T. McKee (fig. 17); T. — capitulata. 23839 capitulata: BLUMEA 340 VOL. 23, No. 2, 1977 canal is present with an opening into each locule just above the placenta (figs. pubescent residual per locule in an 14), with (fig. (fig. arc the 12) raphe all 2-layered; below the placentas centas and composed Fertile seeds (fig. teguments; contents; inner layer of outer outer i, plane 2 or per outer heavily thickened erect, outer the outer glands 17) the splits through the fruit (figs. 16, and and the present in of each locule outer walls (fig. 16). in- and brown several prismatic crystals per cell; and with the inner and and the crushed layers inner angle layer of inner integument colourless. inner and inner walls respectively. oil the inner integument and the (fig. 19) derived from both tangentially integument outer strongly cordate and overlapping hypocotyl sheath wanting; each mature thickened one to flattened moderately thickened; (fig. 21), the 3 —15 the placenta, anatropous longitudinal testa with integument integument brown, walls of each cell broad, (fig. 15) the In locule, winged, integument with outer on xi), distinctive central column terminated by the pla- a Sterile seeds (fig. 20) consisting of the inner integument 13) 10, 10); ovules the vascular strand and of the ovule potentially fertile. result in of layer of inner between winged of axial flower tissue and the inner o, layer outer circle (T. glauca, fig. the rim of the hypanthium; septae 18) a transverse ovules exserted beyond capsule or strongly in the median integument; nucellus of the flower free within the stylar canal (fig. apex of the outer remnants of the integument with Embryo (fig. 22) straight; cotyledons partly enfolding each other (fig. 23); embryo. DISCUSSION The placing of (Dawson, in a the Tristania Metrosiderinae is subtribe 1976) single ring in or a group which the New Caledonian species to questionable as number of respects arc on the placenta, — the group differs from belong typical in the Metrosideros alternate leaves, fascicled stamens, few ovules distinctively winged ovules and seeds, column in the fruit, and the broad, cordate, the central overlapping cotyledons. REFERENCES Dawson, Wilson, J. W. P. G. of N.S.W. 1976. Blumea 1971. Studies Sydney. 23: 7 in the — 11. systematics of the genus Tristania R. Br. B.Sc. Honours Thesis. Univ.