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http://www.youtube.com/watch?v=iWaX97p6 y9U The first multicellular organisms evolved in water and their entire lives were designed around an aquatic environment – photosynthesis and reproduction took place in the water. Recall Protista, Algae, Fungi… Over time, some organisms adapted to life in drier environments. These organisms evolved structures to acquire, transport, and conserve water, as well as reproductive cycles that enabled them to survive on land. 1. 2. 3. All life needs a constant supply of water as well as a method to protect the loss of that water (desiccation). Land plants need rigid supports to hold their leaves up to the sun for photosynthesis. Land plants need to transport water up from the roots and food down from the leaves. 4. 5. Land plants must exchange water and carbon dioxide with the environment without losing too much water in the process. Land plants must be able to reproduce in a dry environment that lacks water in which the sperm can swim. Also, young embryos of land plants are in danger of drying out. Includes the mosses, liverworts, and hornworts. Like the algae from which they evolved, the bryophytes have life cycles that involve an alteration of generations between a haploid gametophyte and a diploid sporophyte. Also, like the algae, the need water for reproduction to occur. 1. 2. 3. They lack vascular tissue; therefore, most are only a few centimeters tall. They lack a protective surface covering (cuticle) to keep water from evaporating from their cells. The lack true roots which could absorb and transport water. Instead, they have rhizoids which anchor them to the ground. 4. They have sperm cells that must swim through water to fertilize the eggs; therefore, they must live in areas that are wet for at least part of the year. 1. 2. 3. The haploid gametophye is the dominant, obvious stage. It’s what we call the moss plant. Standing water is needed for sperm to swim from the male gametophyte to the female gametophyte to fertilize the eggs. The diploid sporophyte is small and can grow only with nourishment provided by the gametophyte (the sporophyte is the dependent on the gametophyte). http://www.youtube.com/watch?v=xbSZbd02UEg& feature=g-wl The members of the phylum tracheophyta are “true” land plants because they have evolved ways of freeing themselves from dependence on wet environments. Tracheophytes possess vascular tissues which transport water and food throughout the plant. There are two types of vascular tissue: Xylem: movement of water from roots to all parts of the plant. ◦ Xylem tissue is made up of tracheid cells which transport water and have thick, strong cell walls that strengthen stems and help plants stand up (the phylum tracheophyta gets its name from this cell type). Phloem: transport of food and nutrients to all parts of the plant Tracheophytes have also developed true roots and true leaves. True roots absorb and transport water from the soil, and true leaves are organs of photosynthesis with a waxy protective covering called a cuticle that prevents water loss. Ferns are well-developed tracheophytes with vascular tissues, strong roots, creeping stems called rhizomes, and large leaves called fronds. There are over 11,000 species of ferns alive. Diploid sporophyte is the dominant stage in ferns. Gametophyte is tiny, lacks vascular tissue, and can grow only in moist areas. Sexual reproduction still depends upon the presence of standing water for the sperm to swim to eggs – ferns are seedless vascular plants! Fern sporophytes produce haploid spores in sporangia on the underside of their fronds. Sporangia are grouped into larger clusters called sori. (fig 21-11 pg 458 in text). Note – a vascular plant is any plant that has connective tissue (xylem/phloem) http://www.youtube.com/watch?v=c4YtOT0Z 6Ek