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Ch. 24 Notes: Plant Structure & Function Characteristics of Plants Multicellular, Eukaryotic, and Photosynthetic autotrophs Share many characteristics with their green algae ancestors: Have chloroplasts with chl.a, chl. b, and carotenoids Cell walls containing cellulose Store food as starch in plastids Plants have evolved complex bodies with cell specialization for different functions. New adaptations/structures were needed to help with the move to land: 1) Transitionfrom Aquatic Ancestors to Land by Bryophytes: Nonvascular, small (mosses, hornworts, and liverworts) 475 mya NO true roots, leaves, or stems A new mode of reproduction was necessary: Embryophyte = embryo is retained inside female’s ovule for protection and to prevent dessication Alternation of Generations = a haploid gametophyte generation produces and alternates with a diploid sporophyte generation. The sporophyte, in turn, produces gametophytes. 2) Seedless Vascular Plants: Ferns and Horsetails 400 mya Regional specializations of the body developed to extract the resources needed for photosynthesis from the terrestrial environment. o Roots, Shoots, and Leaves (division of labor) Vascular Tissue = special tissues for the (long-distance) transport of water and nutrients through the plant body (between leaves and roots) Lignin = added to cell walls of “woody” plants to help them stand up in the terrestrial environment 3) Origin of the Seed: Gymnosperms 360 mya Seed = plant embryo packaged with a store of food within a resistant coat Gymnosperms = seeds as naked structures o Include the Conifers, Cycads, Ginkgoes, and Gnetophytes 4) Emergence of Flowering Plants: Angiosperms 130 mya Flowers evolved attract pollinators for better fitness Unlike gymnosperms, they bear seeds within the flower’s protective ovules (ovaries) ANGIOSPERM BODY PLAN Each system depends on the others for survival of the whole plant Roots Anchors plants Absorbs and conduct water and nutrients Stores food Produce hormones to regulate growth Absorption of water is greatly enhanced by root hairs = projections of epidermal cells o Increase the surface area of the root. o Ex. Rye plants 14 billion root hair cells = 10,626 km! Water and mineral absorption are also enhanced by mycorrhizae = symbiotic associations between roots and fungi Root nodules = symbiotic associations of plant roots and bacteria capable of fixing nitrogen Stems Axillary bud = An embryonic side shoot found in the angle formed by each leaf and the stem; usually dormant o When triggered, can produce a new branch Leaves Main site of photosynthetis Tissue Systems Continuous throughout the plant 1) Epidermal Tissue System Single layer of tightly packed cells covering and protecting the young parts of the plant o Includes Root Hairs o Aerial parts covered by the waxy cuticle o Trichomes = protect plant from too much sun and may help protect plant from herbivores by making a toxic substance Found on stems, leaves, and reproductive organs Look like “hairs” or small thorns Density of these increases on plants as a function of herbivory 2) Ground Tissue System Occupies the space between epidermal and vascular tissue systems Function in photosynthesis, storage, and support 3) Vascular Tissue System Xylem = transports water and mineral from roots to leaves o Made of Tracheids and Vessel Elements Both are dead at functional maturity o Also function in support Phloem = transports sugar and other organic compounds o Sieve-Tube Members = Chains of phloem cells Cells are alive at functional maturity. o Lack a nucleus, ribosomes, and a distinct vacuole. At least one Companion Cell is connected to each sieve-tube member by many plasmodesmata o The companion cell’s nucleus and ribosomes also serve the sieve-tubes member, which lacks these organelles. o Helps load sugar produced in the mesophyll into sieve-tubes of leaves of some plants. PLANT GROWTH Begins with germination of the seed and continues for the lifespan of the plant. Indeterminate growth is made possible by meristems = perpetually embryonic tissues Primary Growth = elongation; increase in height/length Secondary Growth = increase in girth (circumference) Tissue Organization of Leaves Cloaked by a waxy cuticle o Protects against physical damage, pathogens, and water loss Stomata are pores flanked by guard cells which regulate gas exchange with the surrounding air and photosynthetic cells inside the leaf o Allow for transpiration (water loss from the plant by evaporation) o More on the bottom surface of leaf to minimize water loss Vein = vascular bundles with xylem and phloem o May be wrapped in bundle sheath cells in C4 plants