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Mollusks Mollusks  Huge phylum, wide variety of shapes/sizes  ~85,000 different species  Soft-bodied invertebrates  Have bilateral symmetry  Usually have one or two shells with organs in a fluid filled cavity (shells are internal or external)  Most live in water  Fossils are 500+ million years old Why are Mollusks in the same phylum?  Share similar developmental stages  Many of their lifecycles begin with a trochophore (free-swimming larval stage) Classification of Mollusks  Classified into three common groups based on shell (presence & type) and foot type  Gastropods  Bivalves  Cephalopods Class Gastropoda  Includes conchs, snails, slugs, nudibranchs, limpets  Largest group of mollusks  Shell-less or single shelled  Use a radula (a tongue-like organ with rows of teeth) to get food  Move by a muscular foot on the ventral side  Have foot glands that secrete a layer of mucus for sliding Gastropod Protection  Some can pull inside a single shell when threatened  some also have a hard disk (operculum) on their foot that forms a protective door when they withdraw  Some of don’t have a shell  Slugs hide during the day from predators  Nudibranchs can have poison & are brightly coloured  Can reuse nematocysts from cnidarians they eat! Class Bivalva  Includes clams, oysters, mussels, and scallops  Have a hinged, two-part shell  held together by 1-2 powerful muscles  To open or close their shell they either contract or relax their muscles Bivalves are well adapted for water  Clams can burrow in sand  Mussels attach themselves to a solid surface  Scallops escape predators by rapidly opening and closing their shell Class Cephalopoda  Most specialized and complex mollusks.  Include squid, octopi, cuttlefishes and nautiluses.  Soft-bodied with a head attached to a single foot (foot is divided into tentacles) Cephalopods  Have a well developed head and many tentacles for capturing prey  Eyes that distinguish shape!  Closed circulatory system  Moves blood through the body in a series of closed vessels like humans.  Use jet propulsion to move at speeds of 6 m/s  Water flows over the gills into the mantle  Squeezed out through the siphon Class Polyplacophora  Chitons  Shells made of 8 overlapping plates  Plates are embedded in the tough muscular girdle that surrounds the body  Can hold onto irregular surfaces or roll into a protective ball if dislodged Class Scaphopoda  Tusk shells  Example: dentalium  Used by First Nations as a form of currency Mollusks’ Body Plan  4 parts  1. foot: crawling, burrowing, capturing prey (tentacles)  2. mantle: tissue covering the body  3. shell: secreted by mantle – calcium carbonate  4. visceral mass: internal organs Mollusks’ Body Plan  Mantle  Thin layer of tissue that covers the body organs  Mantle cavity (between soft body and mantle)  Contains the gills that are used to breathe by exchanging oxygen and carbon dioxide in the water  Open Circulatory System  Most mollusk have this  Moves blood through vessels and into open spaces around body organs Mollusk Feeding  Snails & slugs: radula (scrape algae or drill holes)  Bivalves: filter-feeders  Water enters through siphon, passes over gills, food gets caught on mucus, water exits through siphon  Octopi & sea slugs: sharp jaws  Respiration  Aquatic mollusks = gills  Land mollusks = through skin  Excretion  Nephridia remove ammonia from blood Mollusk Circulation  Slow-moving mollusks = open circulatory system  Fast-moving mollusks = closed circulatory system Response: Nervous system?  Bivalves: small ganglia, nerve cords, eyespots  Octopi: brain – memory and complex behaviour Movement  Slugs & Snails: secrete mucus & muscular contractions  Bivalves: foot  Octopi: jet propulsion Reproduction  Gastropods & bivalves: sexual reproduction – external fertilization  Cephalopods: internal fertilization Ecology of Mollusks  Food web:  eat plants, animals, filter algae & eat detritus  food source (including us!)  Monitor water quality: filter-feeders concentrate dangerous pollutants  Early environmental warning system  Cancer research? – mollusks don’t seem to develop caner Invasive, non-native species  Zebra mussels: introduced to North America from Eastern Europe & Asia on boats in the 1980s  Spread through Great Lakes and into rivers  Few natural predators  Reproduce rapidly  Attach to anything & in layers Clam Gardens  Made by First Nations from Alaska to Washington  Expand clams’ specific intertidal habitat to increase their growth Clam Gardens  Rock-walled beach terraces  Extend the beach flat  Maintenance:  Constantly dig to allow oxygen water in  Remove shifted boulders