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CRANIAL NERVES - LECTURE B NBIO 401 – Friday, October 8, 2009 1 Abducens nucleus Abducens nerve 2 Genu of the Facial nerve Abducens nucleus Facial nucleus Facial nerve 3 Facial nerve fibers (looping around the abducens nucleus) Facial motor nucleus Abducens Nucleus (eye muscle motoneurons) Facial motor nucleus (muscles of facial expression) 4 Abducens nucleus Solitary nucleus (outside) Superior salivatory nucleus Facial intermediate nerve Spinal trigeminal nucleus Solitary tract (inside) Facial intermediate nerve part of facial nerve carrying: Motor 1) motor output from the superior salivatory nucleus to some salivary glands (GVE) Sensory 1) taste information from the anterior 2/3 of the tongue to solitary nucleus via the solitary tract (SVA) 2) touch information from a small patch of skin on the ear and behind the ear to spinal trigeminal nucleus (GSA) 5 Superior cerebellar peduncle Spinocerebellar fibers (on route to the cerebellum) Inferior cerebellar peduncle Middle cerebellar peduncle Superior cochlear nucleus Vestibulocochlear Nerve VIII Cochlear component (carries auditory signals from the cochlea to the cochlear nucleus) 6 Inferior vestibular nucleus Medial vestibular nucleus Inferior Cerebellar peduncle Vestibulocochlear Nerve VIII vestibular component (carries vestibular signals from the vestibular labyrinth to the vestibular nuclei) 7 Superior Cochlear nucleus Vestibulocochlear Nerve VIII Vestibular nuclei 8 Vestibular nuclei Spinal trigeminal nucleus & tract Hypoglossal Nucleus MLF Solitary tract & nucleus Inferior cerebellar peduncle (spinocerebellar fibers to cerebellum) Cochlear Nucleus (auditory information) Spinothalamic Tract Medial lemniscus Inferior olive nucleus Pyramidal Tract Abducens nerve (to eye muscle) Pontine Gray matter (relay signals to cerebellum) 9 Inferior salivatory nucleus Solitary nucleus & tract Spinal trigeminal Nucleus Nucleus Ambiguus (motoneurons for larynx & pharynx muscles) Glossopharyngeal nerve IX carries: Motor 1) signals to the pharynx from nucleus ambiguus 2) motor output from the inferior salivatory nucleus to some salivary glands Sensory 1) taste information from posterior 1/3 of tongue to the solitary nucleus via the solitary tract. 2) touch information from the posterior 1/3 of the tongue, pharynx, some skin around the ear, internal surface of ear drum to the spinal trigeminal tract 3) Pressure signals from the carotid artery 10 Dorsal motor nucleus of the Vagus Solitary nucleus & tract Nucleus ambiguus Spinal trigeminal nucleus Roots of the vagus nerve X carries: Motor 1) motor signals to larynx & pharynx muscles from motoneurons in nucleus ambiguus. 2) parasympathetic output to the larynx & pharynx & to organs of the chest & abdomen from the dorsal motor nucleus of the vagus Sensory 1) touch signals from pharynx, external ear canal, external side of the ear drum, & skin of the ear to the spinal trigeminal nucleus. 2) taste information from the epiglottus to the solitary nucleus 3) sensory signals from the larynx, trachea, esophagus, chest & abdomen viscera, & pressure & chemoreceptors in the aorta to the caudal part of the solitary nucleus. 11 Nucleus Ambiguus Lateral (motoneurons for larynx cuneate & pharynx muscles) nucleus Vestibular Hypoglossal nuclei Nucleus (tongue motoneurons) Solitary tract & nucleus DMX MLF Spinal trigeminal nucleus & tract Spinocerebellar Tract Spinothalamic Tract Vagus Nerve Rootlet (motor commands to viscera in body) Medial lemniscus (touch information from body) Hypoglossal nerve (motor commands to tongue) Pyramids Inferior olive nucleus (big input to cerebellum) 12 Spinal accessory nerve Spinal accessory roots Spinal accessory nucleus (motoneurons for sternocleidomastoid & trapezius muscles) 13 Gracile fasciculus Gracile Nucleus Cuneate nucleus Cuneate fasciculus Spinal trigeminal nucleus & tract Spinal Accessory Nucleus MLF Pyramidal Decussation Pyramidal tract Spinothalamic Tract 14 Vestibular nuclei Solitary nucleus & tract Dorsal motor nuclei of the vagus External cuneate nucleus Spinal trigeminal nucleus & tract Nucleus ambiguus Hypoglossal nuclei (motoneuron for tongue muscles) Inferior olive Hypoglossal nerve Pyramidal tract 15 Vestibular nuclei Lateral cuneate nucleus Hypoglossal Nucleus Solitary tract & nucleus Spinal trigeminal nucleus & tract DMX IVth ventricle Inferior olive Pyramidal tract Medial lemniscus 16 Somatic Motor Nuclei Oculomotor nucleus Hypoglossal nucleus Oculomotor nerve Trochlear nerve Hypoglossal nerve Trochlear nucleus Abducens nerve Abducens nucleus 17 Special visceral (Branchial) Motor Nuclei Trigeminal motor nucleus Trigeminal nerve Facial motor nerve Facial motor nucleus Nucleus ambiguus axons bound for glossopharyngeal & vagus nerves Nucleus ambiguus 18 General Visceral Motor Nuclei Dorsal motor nucleus of the vagus Axons from the dorsal motor nucleus of the vagus bound for the vagus nerve Inferior salivatory nucleus Superior salivatory nucleus Axons from the superior salivatory nucleus bound for the facial nerve Axons from the inferior salivatory nucleus bound for the glossopharyngeal nerve Edinger-Westphal nucleus Edinger-Westphal fibers bound for the oculomotor nerve 19 Somatic Sensory Nuclei Main sensory trigeminal nucleus Mesencephalic Mesencephalic trigeminal tracttrigeminal nucleus Trigeminal nerve (touch, conscious& unconscious proprioception, pain & temperature from the head to main sensory trigeminal nucleus, mesencephalic trigeminal nucleus & spinal trigeminal nucleus) Spinal Facial intermediate nerve Glossopharyngeal Vagus nerve Spinal trigeminal (touch from ear to spinal Nerve (touch from ear trigeminal nerve trigeminal nucleus) (touch from ear & pharynx & pharynx to nucleus to spinal trigeminal) spinal trigeminal) 20 Visceral Sensory Nuclei Facial intermediate Nerve (taste from anterior 2/3 of tongue to solitary nucleus) Glossopharyngeal nerve (taste from posterior 1/3 of tongue to solitary nucleus) Vagus nerve (taste from epiglottus to rostral solitary nucleus, signals from the trachea, esophagus, chest, abdomen & aorta to caudal part of solitary nucleus) SOLITARY NUCLEUS Rostral 1/2 (taste) TONGUE GENICULATE GANGLION VII - FACIAL (facial intermediate nerve) IX - GLOSSOPHARYNGEAL ANTERIOR 2/3 PETROSAL GANGLION POSTERIOR 1/3 (SVA) EPIGLOTTIS X - VAGUS Caudal 1/2 (autonomic sensory processing) (GVA) NODOSE GANGLION