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E&E Some evidence for evolution A. Direct Observation of Microevolution • Adaptation and Speciation • Natural Selection (many examples, a few covered previously) B. The Fossil Record - gives us much information • stratification of fossils • dating of fossils, sediments – radiometric dating – fission track dating – electron spin resonance dating – thermoluminescence dating Sedimentary Layers – The Grand Canyon Fossil Crinoid Stems and Brachiopods A Trilobite (Odontochile formosa) from Lower Devonian of the Kinglake district (Source: Museum Victoria.) (Trilobites lived from the Cambrian – Permian; 545 – 251 mya) Geologic Time Scale Fission Track Dating of Uranium The fossil record yields… • transitional/intermediate forms, vertebrate examples (see http://www.talkorigins.org/faqs/faq-transitional.html for indepth treatment). • pageant of whale (e.g. transitional Ambulocetus), and horse evolution (Illustrated on the following slides.) • many hominid transitions (e.g. Australopithecus is transitional between Pan and Homo – we’ll study later) • Archaeopteryx, Confuciousornis, Protopteryx are transitional between reptiles and birds (Illustrated in a few slides.) • Eusthenopteron (fish) is linked to the early amphibian Icthyostega; Acanthostega • Genera such as Probainognathus, Probelesodon and Massetognathus (cynodonts) are transitional between reptiles and mammals Biogeographical Evidence coupled to fossil record hummingbirds marsupials Quick note on the phenotype Vp = Vg + Ve + Vg e x g e p A∙MX B∙MY A∙MX A∙NY Gradual versus punctuated evolution • Gradual change over time • Branching, punctuated change Mesonychid – shared ancestry with whales Ambulocetus Rodhocetus, a predominantly aquatic whale from the Eocene (47 mya) Vestigial pelvis and femur in modern whales Many dinosaur bones have been found . . . (Ultrasaurus) Archaeopteryx Comparative anatomy Homology having common ancestry but not necessarily retaining similarity of structure or function The Adult Examples: • Mammalian forelimbs (“hands”) o grasping hands of primates o digging hands of moles o locomotive hands of horses o paddles of porpoises o wings of bats • Vertebrate hindlimb structure Hind Limb Structure in Vertebrates Homo sapiens The Embryo Bio-molecular Evidence •Protein evidence •Nucleic acid evidence •Others… RELATIVE LEVEL OF SHARED CHARACTERISTICS AMONG SPECIES DNA Phylogeny EVIDENCE FOR EVOLUTION 1. Fossil Record 2. Comparative Anatomy (phenotype) physiology, anatomy, behavior, etc… 3. Biogeographical 4. Biochemical